Paduniella nama, Johanson, Kjell Arne & Oláh, János, 2010

Johanson, Kjell Arne & Oláh, János, 2010, Description of six new species of Oriental Paduniella (Insecta: Trichoptera: Psychomyiidae), Zootaxa 2548, pp. 43-56 : 51-52

Paduniella nama , new species

Figs 16–18

Paduniella nama , new species is unique in the combination of the shape of the inferior appendages, presence of a triangular process at basis of each of the inferior appendages, long and straight superior appendages, and a visible median branch and ventral lobe of the inferior appendages. The phallic apparatus is unique in having a relatively short median process and in the shape of the phallicata apex. The genitalia resemble those of P. semarangensis Ulmer ( Thailand) , P. hatyaiensis Malicky & Chantaramongkol and P. nike Malicky ( Nepal) . Paduniella nama is separated from P. semarangensis and P. n i k e in the shape of the phallic apparatus, particularly in possessing shorter median process and in the shape of the phallicata; and from hatyaiensis particularly by the presence of two protuberance at dorsal and ventral corners of the posterior margin of the ventral part of sternite IX.

Male genitalia. Sternite IX narrowly club-shaped in lateral view ( Fig. 16 ); wide ventral part with sigmoid anterior margin and straight posterior margin with small protuberance at dorsal and ventral corners; dorsal part of sternite IX tapering dorsad along its length, clearly more slender than ventral part, sigmoid, as long as ventral part, orienting anterodorsad before curving dorsad at dorsal apex; joining tergites IX well above midheight of genitalia. Narrow dorsal part of sternite IX forming right angle with ventral part of sternite IX. Dorsal apex of sternite IX connecting to sternal strip near dorsal apex of strip ( Fig. 18 ). Sclerotized part of tergite IX long, boomerang-shaped in lateral view ( Fig. 16 ), basal half orienting posteroventrally, bending 90 degrees dorsad before weakly curving posterodorsally subapically; slightly exceeding membranous part above superior appendages; membranous part covered by long microtrichia. Superior appendages long, dorsal and ventral margins nearly parallel-sided in lateral view; dorsal margin straight, with few setae; ventral margin slightly undulating; apex rounded. Median branch of inferior appendages visible above main branch of inferior appendage in lateral view ( Fig. 16 ), in ventral view ( Fig. 17 ), median branches forming slightly convergent processes with irregular apices; originating before mid-length of main branch ( Fig. 16 ). Ventral lobe of inferior appendages with ventral margin parallel with ventral margin of main branch in lateral view ( Fig. 16 ); with ventral setae; in ventral view ( Fig. 17 ) slender, digitate, straight, orienting posterad. Additional process present basiventrally on main branch, sharply triangular in lateral view ( Fig. 16 ); digitate in ventral view; covered by setae. Main branch of inferior appendages straight before slightly curving dorsad at apex; dorsal margin nearly straight, interrupted by dorsal margin of median branch; ventral margin irregularly convex ( Fig. 16 ); apex truncate, except slightly produced posterad at dorsal corner; in ventral view ( Fig. 17 ), main branch tapering along its length, with smooth, triangular mesal corner. Phallic apparatus connecting to sternite IX and tergite IX by long, slender, nearly parallel-sided sternal strip ( Fig. 18 ). Phallobase narrow, shorter than sternal strip; oriented anterodorsally before curving anterad at apex. Median process about as long as height of sternite IX; bow-shaped, parallel-sided along its length. Anterior 3/5ths of phallicata narrow, parallel-sided; bow-shaped; about 2 times wider than median process; widening into asymmetrical clubshaped apex with nearly straight dorsal margin and strongly convex ventral margin; apex rounded. Paramere complicated, apparently as wide as median process, with strongly sclerotized dorsal and ventral margins, membranous in between; nearly parallel with dorsal margin of phallicata, ending into pointed apex; as long as phallicata.

Male holotype: VIETNAM, Ngoc Lac, 25–26.i.1986, light, leg. János Oláh (OPC, alcohol).

Paratypes: same data as for holotype— 2 males (OPC, alcohol); same data as for holotype— 2 males (NRM, alcohol); VIETNAM, Hoabinh Hotel, 23.x.1986, light, leg. János Oláh— 2 males (OPC, alcohol).

Distribution: Vietnam.

Etymology: Nama , from Vietnamese “ nama ”, meaning five, referring to this species being the fifth Paduniella species from Vietnam treated in this paper.

Figures

FIGURES 16 – 18. Paduniella nama, new species, male holotype. 16 — genitalia, lateral; 17 — genitalia, ventral; 18 — phallic apparatus, lateral. Scale bar = 0.2 mm, refers to all figures.