Garra gallagheri Krupp, 1988

Kirchner, Sandra, Kruckenhauser, Luise, Pichler, Arthur, Borkenhagen, Kai & Freyhof, Jörg, 2020, Revision of the Garra species of the Hajar Mountains in Oman and the United Arab Emirates with the description of two new species (Teleostei: Cyprinidae), Zootaxa 4751 (3), pp. 521-545 : 528-529

Garra gallagheri Krupp, 1988

( Fig. 4 )

Garra barreimiae gallagheri, Krupp, 1988:402 (type locality: Wadi Bani Khalid north of Muqal, 22°40’N, 59°05’E, Oman); Garra barreimiae Gallagheri Clade (Kirchner et al. submitted).

Material examined. FSJF 3536 , 20 , 46–64 mm SL; Oman: Wadi Bani Khalid at Sayh al Hayl, 22.595°N, 59.086°E .— FSJF 4094 , 11 , 47–61 mm SL; Oman: Wadi Bani Khalid north of Muqal, 22.619°N, 59.093°E .— FSJF 4090 , 2 , 41–51 mm SL; Oman: Falaj in Bani Bu Ali , 22.023°N, 59.317°E .— FSJF 4089 , 6 , 34–38 mm SL; Oman: Falaj in Al Mudayrib , 22.612°N, 58.667°E .

Material used in molecular genetic analysis. FSJF DNA-3248 ; Oman: Falaj in Bani Bu Ali , 22.023°N, 59.317°E. (GenBank accession numbers: MN830869 , MN830870 ).—NMW-100030; NMW-100042_1; NMW- 100042_2 , FSJF DNA-2578 ; Oman: Wadi Bani Khalid at Sayh al Hayl, 22.595°N, 59.086°E (GenBank accession numbers: MN830409 , MN830410 , MN830411 , KX244645 , KX244646 , KX244647 , KX244648 ) .— FSJF DNA- 3247 ; Oman: Falaj in Al Mudayrib , 22.612°N, 58.667°E. (GenBank accession number: MN 830871 ) .

Diagnosis. Garra gallagheri differs from G. barreimiae by having a plain or almost plain, greenish or black flank pattern (vs. strongly mottled), with a faint dark-green lateral stripe present in some individuals (vs. absent), orange midlateral scales on the flank absent (vs. present), orange spot at the upper opercle absent (vs. present), dorsal-fin tip not white (vs. white), and 12–14 gill rakers on the lower limb of the first gill arch (vs. 15–18).

Distribution. Garra gallagheri is endemic to Oman, where it is distributed in the Wadi Bani Khalid and isolated falaj irrigation systems downstream, for example in the cities Al Mudayrib and Bani Bu Ali ( Fig. 2 ).

Remarks. When Krupp (1988) described G. b. gallagheri he did not mention G. longipinnis , as this species was believed to be characterized by its very large fins (see below). Krupp also did not mention whether he examined any comparative material of G. barreimiae . He states that G. barreimiae is distributed in the coastal drainages of Oman, but this statement seems to be based on the work of Banister & Clarke (1977). Krupp distinguished G. b. gallagheri from G. b. barreimiae by possession of 8 dorsal-fin rays (vs. 7). It is likely that Krupp counted the posterior pair of fin rays based on one pterygophore (here counted as 11 / 2 rays) as two rays, thus 8 rays as counted by Krupp (1988) can be treated as 7½ rays. All 20 individuals of G. gallagheri from the type locality examined in the present study possessed 7½ branched dorsal-fin rays.

Figures

FIGURE 2. Records of Garra species from the Hajar Mountains based on material analysed in this study.

FIGURE 4. Garra gallagheri, from top: FSJF 3536, 60, mm SL; FSJF 4094, 55 mm SL; 62 mm SL; Wadi Bani Khalid, Oman.