Anthrax dentata Becker, 1906

Yao, Gang & Yang, Ding, 2018, Anthrax, newly recorded from Vietnam (Diptera: Bombyliidae), Zoological Systematics 43 (2), pp. 227-230 : 228-229

Anthrax dentata Becker, 1906 ( Figs 1–7 )

Argyromoeba dentata Becker, 1906: 148 . Type locality: Tunisia (1 syntype in ZMHB).

Anthrax efflatouni Paramonov, 1935: 7 , 22 (key only); 1936: 90. Type locality: Egypt (Holotype (destroyed) in PPDD).

Diagnosis. Occiput with sparse black hairs and white scales, densely covered erect brown hairs on margin; mesonotum with long dense white hairs at anterior margin; thorax with anepisternum and katepisternum with long brown hairs and white pollen, wing with join of vein R 4 and R 5 and join of vein dm-cu and CuA with brown spots; tergites 5–7 of abdomen with dense white scales laterally.

Redescription. Male. Body length 7 mm, wing length 9 mm.

Head black with white pollen except mid frons with brown pollen. Hairs on head mostly black, scales white; ocellar tubercle reddish with sparse black hairs, frons narrowing directed apically, with black erect hairs and white recumbent scales, face with black erect hairs and white recumbent scales; occiput with sparse black hairs and white scales, and with dense erect brown hairs on edge. Antenna black with white pollen; scape thick with long black hairs; pedicel slightly wider than long, with short black hairs; first flagellomere onion-shaped, bare, tip with tufty brown hairs. Antennal ratio: 2:1:2. Proboscis brown with yellow hairs.

Thorax black with pale pollen, scales black and white. Hairs on thorax black and white; postpronotal lobe with long dense black hairs, mesonotum with long dense white hairs at anterior margin; thorax with sparse black and white scales on back, anepisternum and katepisternum with long brown hairs and white pollen. Scutellum black with pale pollen, with six scutellar bristles on posterior edge. Legs brown except fore tarsi black. Hairs on legs mostly black, bristles black, scales yellow. Femora with dense yellow scales; tibiae and tarsi with short sparse black hairs. Mid femur with six av; hind femur nine av. Mid tibia with seven ad, 11 pd, seven av, and seven pv; hind tibia with nine ad, nine pd, seven av, and six pv. Wing ( Fig. 1 ) half brown half hyaline, joint of vein R 4 and R 5 and joint of vein dm-cu and CuA with brown spots. Bend of vein R 4 with appendix, vein r-m nearly at middle of cell dm. Base of vein C with brush-like long black bristles. Halteres brown.

Abdomen black with white pollen. Hairs on abdomen white. Tergite 1 with white erect hairs and white recumbent scales; tergites 2–4 bare, tergites 5–7 with dense white scales laterally. Sternites brown with black erect hairs and white recumbent scales.

Male genitalia ( Figs 2–7 ). Epandrium subtriangular, distinctly higher than long, cercus well exposed in lateral view; epandrium trapezoidal, with lateral extension at base in dorsal view; gonocoxite sharply narrowing at middle in ventral view, extended apically; gonostylus elongate, its obtuse tip slightly curved in lateral view; epiphallus turriform, tip mushroom shaped in dorsal view, with an acute tip in lateral view.

Female. Body length 11 mm, wing length 14 mm. Similar to male, but mesonotum with long white and black hairs at anterior margin; tergites 5–7 bare.

Material examined. 1 male, Vietnam, Dak LaK, Yok Don National Park , 8.V.2012, Xingyue Liu ; 1 female, Vietnam, Bac kan, Ba Be National Park , 21.V.2012, Xingyue Liu .

Distribution. Vietnam (Dak LaK).

Remarks. The species identified as A. dentata , is belongs to Anthrax dentata complex, similar to Anthrax trifasciatus complex, In the A. trifasciatus complex, the wing is partly infuscated, with the infuscated part no more that the basal half of the anal and axillary cells, and there is a broad clear area basal to the r-m cross-vein often extending forward to R 1 and the basal cells often have extensive clear areas in the middle; In the A. dentata complex, the wing is partly infuscated, the infuscated part tends to be more extensive and uniform, and reaching to at least the middle of the anal and axillary cells, without or with only a narrow clear area basal to the r-m crossvein and without extensive clear areas in the centres of the basal cells.

Funding This research is supported by the National Natural Science Foundation of China (31301670) and China Postdoctoral Science Foundation (2015M581205).

Acknowledgements We are very grateful to Dr. Xingyue Liu from CAU for collecting the specimens. Our thanks also go to Dr. Jeff Skevington from CNC for providing photos. We also heartily to thank Dr. Christine Lambink from Queensland Museum for providing really helpful suggestions and references. Two reviewers are also thanked for providing useful comments on an earlier draft of this paper.

Figures

Figure 1. Anthrax dentata Becker, 1906, wing.

Figures 2–7. Anthrax dentata Becker, male genitalia. 2. Epandrium and cercus, dorsal view. 3. Epandrium and cercus, lateral view. 4. Phallus, lateral view. 5. Gonocoxa and gonostylus, lateral view. 6. Gonocoxa and gonostylus, ventral view. 7. Phallus, dorsal view.