Yoruba, Rodrigues & Rheims, 2020

Rodrigues, Bruno V. B. & Rheims, Cristina A., 2020, An overview of the African genera of Prodidominae spiders: descriptions and remarks (Araneae: Gnaphosidae), Zootaxa 4799 (1), pp. 1-80 : 69

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gen. nov.

Yoruba gen. nov.

Etymology. The generic name is a noun in apposition that refers to Yoruba language, one of the official languages of Nigeria and widely spoken in western Africa, where the new genus was recorded. The gender is masculine.

Type species. Yoruba ibadanus sp. nov.

Diagnosis. Species of Yoruba gen. nov. are distinguished from those of other Prodidominae genera by male palp with a ventral tibial apophysis distinctly separate from the retrolateral tibial apophysis ( Figs 58D View FIGURE 58 , 59B View FIGURE 59 , 60B View FIGURE 60 ), embolus robust with a membranous base ( Figs 59B View FIGURE 59 , 60A View FIGURE 60 ), and long and laminar conductor with the terminal part sclerotized ( Figs 59B View FIGURE 59 , 60A View FIGURE 60 ); female epigyne with wide copulatory openings located medially ( Figs 59D View FIGURE 59 , 60E View FIGURE 60 ); vulva with copulatory ducts enlarged, with small secondary spermathecae connected, connecting duct slender and convoluted laterally, close to primary spermathecae ( Figs 59E View FIGURE 59 , 60F View FIGURE 60 ).

Description. Total length males 1.97–2.40 and females 2.12–2.7. Carapace and legs pale yellow, abdomen slightly gray ( Figs 54A, B View FIGURE 54 ). Carapace longer than wide, slightly narrowed at cephalic region, oval ( Figs 54A, B View FIGURE 54 ). Fovea absent. Eight eyes, posterior row strongly procurved and anterior row approximately straight ( Figs 54C View FIGURE 54 , 55A View FIGURE 55 ); PME and PLE irregular; AME dark; interdistance contiguous of AME–ALE, PME–PLE, ALE–PLE. Chilum absent. Chelicerae relatively small (0.22–0.27), without boss or teeth ( Figs 55D, E View FIGURE 55 ); fang with shaft serrula. Endites slightly convergent anteriorly ( Figs 54D View FIGURE 54 , 55B View FIGURE 55 ), with few hairs on internal margin ( Fig. 55C View FIGURE 55 ); serrula absent; labium longer than wide ( Figs 54D View FIGURE 54 , 55B View FIGURE 55 ). Sternum longer than wide, anterior margin straight, rebordered anteriorly and laterally ( Fig. 54D View FIGURE 54 ); posterior region strongly protruding between coxae IV, with numerous long and erect setae; intercoxal setae absent and precoxal triangles present ( Fig. 54D View FIGURE 54 ). Pedicel with ventral sclerite triangular. Leg formula 4123. Femur I and II with dorsal spines; legs III and IV with spines on femora, tibiae and metatarsi. Patella I larger than the others, length similar to tibia I. All tarsi and metatarsi without scopulae. Trichobothria present on dorsal tibiae, metatarsi and tarsi, arranged in two rows; bothrium with 3–4 ridges in proximal plate ( Fig. 56D View FIGURE 56 ). Tarsal organ capsulate, with oval opening, located dorsally and distally on tarsi ( Fig. 56E View FIGURE 56 ). Two smooth claws ( Fig. 56B View FIGURE 56 ); claw tufts composed of slightly widened tenent setae inserted in well-delimited plate; solid claw tuft clasper present ( Figs 56B, C View FIGURE 56 ). Abdomen oval, longer than wide ( Figs 54A, B View FIGURE 54 ), without scales; dorsum of abdomen anteriorly with long, dark, posteriorly curved setae. Six spinnerets; ALS longer than wide, separated from each other by less than their diameter ( Figs 54E, F View FIGURE 54 ), piriform gland spigots cracked and elongated, with associated setae ( Figs 57A, B View FIGURE 57 ), major ampulate gland spigot field on well-defined conical structure ( Fig. 57C View FIGURE 57 ), male with one major ampulate gland spigot and female with two; PLS and PMS conical and short, PLS larger than PMS ( Figs 57A View FIGURE 57 , 58A View FIGURE 58 ); PLS in male apparently with one minor ampulate gland spigot and two aciniform gland spigots ( Fig. 58B View FIGURE 58 ), females apparently with one minor ampulate gland spigot, two aciniform gland spigots and one cylindrical gland spigot ( Fig. 57E View FIGURE 57 ); PMS in male with one minor ampulate gland spigot and one aciniform gland spigot ( Fig. 58C View FIGURE 58 ), females apparently with one minor ampulate gland spigot, one aciniform gland spigot and one cylindrical gland spigot ( Fig. 57F View FIGURE 57 ). Female palp tarsus with chemosensory patch apical and truncad, without claw ( Fig. 56F View FIGURE 56 ). Male palp: femur unmodified, with one dorsal median spines; tibia short, with RTA and VTA ( Figs 58D, E View FIGURE 58 ); cymbium with apical scopula ( Figs 59A View FIGURE 59 , 60C View FIGURE 60 ); bulb ovoid ( Fig. 60A View FIGURE 60 ); sperm duct visible ventrally with terminal part narrow; embolus arising prolaterally ( Figs 58D View FIGURE 58 , 59B View FIGURE 59 , 60A View FIGURE 60 ); conductor with membranous base, slightly twisted and arising medially ( Figs 59B View FIGURE 59 , 60A View FIGURE 60 ). Vulva: enlarged copulatory ducts; secondary spermathecae small and positioned distally on duct; slender connecting ducts; primary spermathecae separated from each other; fertilization ducts laterad ( Figs 59E View FIGURE 59 , 60F View FIGURE 60 ).

Distribution. Nigeria and Ivory Coast ( Fig. 61 View FIGURE 61 ).

Composition. Two species: Yoruba ibadanus sp. nov. and Y. toubensis sp. nov.