Adelognathus leucotrochi Shaw & Wahl

Shaw, Mark R. & Wahl, David B., 2014, Biology, early stages and description of a new species of Adelognathus Holmgren (Hymenoptera: Ichneumonidae: Adelognathinae), Zootaxa 3884 (3), pp. 235-252: 237-240

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Adelognathus leucotrochi Shaw & Wahl

sp. n.

Adelognathus leucotrochi Shaw & Wahl   , sp. n.

( Figs 1–7 View FIGURES 1–3 View FIGURES 4–7 )

Type material: Holotype: UNITED KINGDOM, England, Cumbria, Beetham : ♀ “ Nematus leucotrochus   on Ribes uva-crispum   with 2 ectos, 26.5.1991, em. 21.4.1992; NMSZ 1993.143” (M.R. Shaw) ( RSME). Condition of holotype: intact   . Paratypes: similar data as holotype but often different brood size, collection and emergence dates (collection dates of 24.v.1991, 25.v.1991, 26.v.1991, and 28.v.1991, emergence dates in iv.1992), 36♀, 20♂ ( AEIC, HNHM, NHML, RMNH, RSME, ZINC); UNITED KINGDOM, Scotland, Edinburgh, Grange (suburbs), from same host species, collected and, emerged iv.1999 (M.R. Shaw), 6♀ ( RSME). Non-paratype material: 2♀   collected in England (North Yorkshire, Tadcaster , 24.iv.2011 (W.A. Ely )) and Scotland (Edinburgh, Grange, 3.v.1990, (M.R. Shaw ))   .

Diagnosis. This species ( Fig. 1 View FIGURES 1–3 ) can be recognized by the following combination of characters: clypeus, supraclypeal area, and paraocular area yellow ( Fig. 5 View FIGURES 4–7 ); clypeal apex truncate; flagellomere 4 about 2.2 × as long as wide; subocular sulcus absent; mesoscutum smooth and evenly setose; mesopleuron centrally glabrous; hind coxa white; hind leg with tarsomere 2 about 1.1 × as long as tarsomere 5; vein 3rs-m of fore wing spectral (and areolet superficially appearing open in many specimens); T1 smooth; T2 smooth and with widely scattered minute setiferous punctures medially and laterally, mediolaterally impunctate.

Description. Female. Structure. Scape about 2.0 × as long as pedicel (measured on ventral surface); flagellum about 0.8 × length of hind wing; 12 flagellomeres, flagellomeres 7–12 distinctly thickened ( Fig. 6 View FIGURES 4–7 ); flagellomere 4 about 2.2 × as long as wide. Clypeus smooth with weak transverse grooves, apex truncate and narrowly reflexed ( Fig. 5 View FIGURES 4–7 ). Labrum extending beyond clypeus, its apical margin distinctly concave ( Fig. 5 View FIGURES 4–7 ). Supraclypeal area granulose. Malar space about 0.3 × as long as basal mandibular width. Subocular sulcus absent. Occipital carina not elevated as lamella. Hypostomal carina joining occipital carina before base of mandible. Propleuron weakly convex. Mesoscutum smooth, with minute setiferous punctures separated by 3.0–5.0 times their diameter ( Fig. 2 View FIGURES 1–3 ); lateral lobes weakly and shallowly foveolate. Epicnemial carina complete, its dorsal end touching anterior margin of mesopleuron. Mesopleuron smooth and centrally glabrous ( Fig. 3 View FIGURES 1–3 ). Propodeum with anterior and posterior transverse carinae absent, anterior 0.5 of median longitudinal carina absent; anterior 0.3–0.5 smooth, posterior 0.5–0.7 with numerous more or less longitudinal rugulae on granulate surface ( Fig. 2 View FIGURES 1–3 ). Tarsomere 4 of fore leg about 1.8 × as long as wide; hind leg ( Fig. 7 View FIGURES 4–7 ) with tarsomere 2 about 1.1 × as long as tarsomere 5. Vein 3rs-m of fore wing spectral (and areolet superficially appearing open in many specimens). T1 1.3–1.5 × as long as apical width, appearing subpetiolate in dorsal view ( Fig. 2 View FIGURES 1–3 ), smooth and shining; S1 about 0.2 × as long as T1 ( Fig. 7 View FIGURES 4–7 ). T2 smooth and with widely scattered minute setiferous punctures medially and laterally (punctures separated by at least 1.0 × setal length), mediolaterally impunctate ( Fig. 4 View FIGURES 4–7 ). Color. Black, the following yellow: mandible except for dark brown apex, gena immediately adjacent to mandible, labrum, clypeus, supraclypeal area, paraocular area (extending 0.6 × distance between antennal socket and apex of eye), ventral surfaces of scape and pedicel, ventral 0.5 of propleuron, ventral 0.7 of anterior edge and dorsal posterior corner of lateral area of pronotum, subalar ridge and immediately adjacent area of mesopleuron, tegula, apical 0.2 of T2, apical 0.4 of T3–4, apical 0.5–0.6 of T5, and following tergites. Flagellum with ventral surface light brown, dorsal surface dark brown. Legs with coxae, trochanters, and trochantelli white, except for diffuse light brown of basal 0.1 of hind coxa; femora, tibiae, and tarsi brownish-yellow. Pterostigma of fore wing light brown. Length. 3.4–3.7 mm (3.6 mm); fore wing 3.7–3.9 mm (3.8 mm).

Male. Structure. As in female, except flagellomeres 6–8 with elevated linear tyloids. Color. As in female, except: yellow of paraocular area extending to 0.8–0.9 × distance between antennal socket and apex of eye; ventral surface of flagellum brownish-yellow; lateral area of pronotum yellow but for brown stripe extending from median area to center of lateral area. Length. 3.1–3.5 mm; fore wing 2.9–3.2 mm.

Comments. In Fitton et al. (1982), the female of A. leucotrochi   runs to couplet 9, where it founders due to the short antenna, and yellow supraclypeal area and paraocular area. In Kasparyan (1990), A. leucotrochi   will go smoothly to couplet 56, which leads to A. brevicornis Holmgren   and A. rufithorax Kasparyan. However   , the combination of the length of flagellomere 4 and color patterns of the supraclypeal area, propleuron, and mesopleuron rules out placement in either species. Comparisons with determined material (including specimens used by Kasparyan) and published descriptions convince us that A. leucotrochi   cannot be construed as belonging to any described species. It can be distinguished by the combination of: distinctly thickened flagellomeres 7–12, flagellomere 4 about 2.2 × as long as wide, color pattern of anterior head surface ( Fig. 5 View FIGURES 4–7 ), regularly punctate mesoscutum (thus appearing evenly setose in pristine specimens), black mesopleuron and propodeum, length of S1 about 0.2 × length of T1, and smooth surface of T2. In particular, female A. leucotrochi   differ from A. brevicornis   , the closest species to it morphologically, in its completely yellow supraclypeal area, more extensively yellow portion of the gena immediately adjacent to the mandible, more extensive yellow markings on the propleuron and pronotum, and almost entirely yellow hind coxa (usually extensively brown in A. brevicornis   ). In males, the pronotal and propleural coloration of the two species is more similar, but in A. leucotrochi   the yellow paraocular area extends further dorsally (about 0.8 × the distance between the antennal socket and the apex of the eye, against 0.5 × in A. brevicornis   ). In addition, A. brevicornis   has a later flight period in Britain (June through July, possibly also August through September) and different hosts (found several times on an undetermined tenthredinid on Betula   ).

Yellow to whitish apical banding is found on the tergites of many species of Adelognathus   . In A. leucotrochi   , the apical bands in tergites 3+ appear to be caused by a particularly thin epidermis, as they are often degraded to the point where a good portion of a band is missing in a mottled and irregular fashion.

Etymology. The specific name derives from that of the host, Nematus leucotrochus Hartig.  


American Entomological Institute


Hungarian Natural History Museum (Termeszettudomanyi Muzeum)


Natural History Museum, Tripoli


National Museum of Natural History, Naturalis