Tetraphlebia C. Felder & R. Felder, 1867

Matz, Jess & Brower, Andrew V. Z., 2016, The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes, Zootaxa 4125 (1), pp. 1-108: 69

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Tetraphlebia C. Felder & R. Felder, 1867


Tetraphlebia C. Felder & R. Felder, 1867 

Type species: T. germainii C. Felder & R. Felder, 1867  = Faunula C. Felder & R. Felder, 1867  syn. nov.

Type species: F. leucoglene C. Felder & R. Felder, 1867 

Diagnosis. Tetraphlebia  has a well-developed M 1 -M 3 ocellus on the ventral side of the forewing with variable pupillation. In T. leucoglene  , males have heavy androconia, but these scales are sparse in T. germainii  and absent in T. eleates  . The hindwing is rectangular with a postmedian band on the ventral side that is without ocelli. Though specimens were unavailable for dissection, T. patagonica (Mabille, 1855)  ( Erebia  ), lately a species of Faunula  (cf. Pyrcz 2012), is very similar to T. leucoglene  in that it bears an identical M 1 -M 3 VFW ocellus with a large, white pupil, but T. patagonica  also bears two ocelli on the VHW, making it the only Tetraphlebia  with hindwing ocelli. Antennae are round in T. eleates  and T. leucoglene  , spatulate in T. germainii  . Eyes are naked and terminal palp segment is short and conical or oval. Foreleg tarsi are segmented with males having 2–3 tarsal segments and females 3–5 segments, those of T. leucoglene  females also bearing spines. Male genitalia with the proximal end of the valvae triangular, a wide pedunculus, an aedeagus that is truncate at the proximal end, a saccus longer than it is wide, and an uncus that is widest where it meets the tegumen, narrowing gradually toward the distal end.

Remarks. Felder & Felder described Faunula  as a new genus immediately following the description of Tetraphlebia  , noting similarities in wing venation between the two genera. These genera are here combined based on morphological and genetic similarity.