Lasioglossum (Dialictus) hudsoniellum ( Cockerell, 1919 )

Gardner, Joel & Gibbs, Jason, 2020, The ‘ red-tailed’ Lasioglossum (Dialictus) (Hymenoptera: Halictidae) of the western Nearctic, European Journal of Taxonomy 725, pp. 1-242 : 90-99

Lasioglossum (Dialictus) hudsoniellum ( Cockerell, 1919)

Figs 1 , 42 , 90D , 107A , 116C

Halictus hudsoniellus Cockerell, 1919: 290 (holotype, ♀, deposited in USNM, examined).

Halictus (Chloralictus) hudsoniellus – Sandhouse 1924: 4 (key).

Lasioglossum (Chloralictus) hudsoniellum – Michener 1951: 1114 (catalog).

Dialictus hudsoniellus – Hurd 1979: 1966 (catalog). — Moure & Hurd 1987: 104 (catalog).

Lasioglossum (Dialictus) hudsoniellum – Gibbs 2010: 140 , figs 116a–c, 117a–g (key, redescription). — Scott et al. 2011: 30 (checklist).

Diagnosis

Females of Lasioglossum hudsoniellum can be recognized by the face shiny with dense but distinctly separated and similarly-sized punctures over the frons and paraocular area, mesepisternum with crowded punctures (i =0 pd), scutum usually moderately densely and evenly punctate (i ≤ 1–2 pd), T1 usually with moderately dense punctures (i=1–2 pd) outlining distinct impunctate apicolateral ovals (present but less distinct in most other species), pronotal angle usually about 90 degrees, postgena shiny, and T2 without dark spiracular spots. They are most similar to L. spivakae sp. nov., L. arenisaltans sp. nov., and L. lilianae sp. nov. Females of L. spivakae sp. nov. have the scutum punctures with interspaces 2–3 pd wide immediately mesad of the parapsidal line, metasoma with tomentum mostly restricted to T3–4, pronotal angle obtuse, and metasomal terga usually with sharply delineated brown to black bands. Females of L. arenisaltans sp. nov. have the frons punctures crowded and indistinct, scutum punctures very dense (i <1 pd), postgena lineate, and T2 with dark spiracular spots. Females of L. lilianae sp. nov. have the scutum and T1 more finely and sparsely punctate (i =1–4 pd in part).

Males of L. hudsoniellum can be recognized by S5 apical rim strongly concave, flagellomeres relatively short (F2 about 1.25 times as long as F1 and about as long as broad), face below eye emargination covered in dense tomentum, T1–3 rims usually moderately densely punctate (i=1–2 pd), and T1–5 with abundant tomentum. They are most similar to L. arenisaltans sp. nov. and L. imbriumbrae sp. nov. Both of these species have S5 apical rim straight and metasoma with tomentum sparse or absent. In addition, males of L. arenisaltans sp. nov. have the flagellomeres relatively longer (F2 about 1.75 times as long as F1 and 1.5 times as long as broad), and males of L. imbriumbrae sp. nov. have T1–3 rims impunctate.

Etymology

Cockerell (1919) named this species after the Hudsonian life zone in the Rocky Mountains, where the type specimen was collected, plus the Latin adjectival diminutive suffix - ellus.

Material examined

Holotype

UNITED STATES – Colorado • ♀; Longs Peak Trail, Hudsonian Zone ; [40.274° N, 105.579° W]; 3200 m a.s.l.; 22 Jul.; Cockerell leg.; USNM 27765 .

Other material

CANADA – Alberta • 3 ♀♀; 11.8 km SSW of Empress (stabilizing dune); 50.8504° N, 110.0489° W; 24 May 2019; T.M. Onuferko leg.; CMNC • 1 ♀; Writing-On-Stone Prov. Pk., Sand-North ; [49.09° N, 111.62° W]; 1 Aug. 1990; M. Klassen leg.; PCYU. – Saskatchewan • 2 ♀♀; N Bitter Lake, Tunstall (stabilizing dune); 50.1456° N, 109.8062° W; 2 Jun. 2019; T.M. Onuferko leg.; CMNC

MEXICO – Baja California Sur • 7 ♀♀; 17 mi. NE of Santa Rita ; [24.76° N, 111.27° W]; 18 Sep. 1983; R.R. Snelling leg.; ex Euphorbia ; LACM • 1 ♀; Constitución , 18 km N; [25.18° N, 111.75° W]; 40 m a.s.l.; 13 Apr. 1994; A. Rodríquez leg.; ex Brassica nigra ; SEMC • 1 ♀; Playa Buenaventura, nr San Buenaventura; [26.64° N, 111.83° W]; 0 m a.s.l.; 14 Apr. 1997; A. Rodríquez leg.; ex Brassica nigra ; SEMC . – Chihuahua • 1 ♀; 3 mi. W of Jimenez ; [27.14° N, 104.97° W]; 1250 m a.s.l.; 26 Jul. 1952; SEMC • 1 ♀; Hwy 45, 5 rd mi. NE of Salaices (15 mi. SW of jct. Hwy 49); [27.08° N, 105.13° W]; 1585 m a.s.l.; 24 Aug. 1976; J.P. and K.E. Donahue leg.; LACM . – Coahuila • 1 ♀; 2 km N of San Lorenzo ; 25.5167° N, 102.1833° W; 1430 m a.s.l.; 24 Mar. 1992; R. Brooks leg.; ex Varilla mexicana ; SEMC • 1 ♀; same location as for preceding; 24 Mar. 1992; B. Alexander leg.; ex Bahia absinthifolia ; SEMC • 1 ♀; 55 km E of Parras de la Fuente ; 25.3167° N, 101.7333° W; 1960 m a.s.l.; 1 Apr. 1992; D. Yanega leg.; ex Dalea eriophylla ; SEMC • 4 ♀♀; Arteaga, San Antonio de las Alazanas ; [25.2847° N, 100.6131° W]; 2015 m a.s.l.; 21 Jul. 2013; LRF and CVM leg.; LRF • 1 ♀; San Antonio de las Alazanas, Arteaga ; 25.2847° N, 100.6131° W; 2015 m a.s.l.; 9 Mar. 2014; LRF and CVM leg.; LRF . – Durango • 2 ♀♀; 36 mi. SW of Cd. Lerdo ; [25.17° N, 103.93° W]; 1113 m a.s.l.; 16 Aug. 1963; Naumann and Willis leg.; SEMC • 1 ♀; Nombre de Dios ; [23.85° N, 104.24° W]; 13 May 1952; F.D. Parker leg.; UCDC . – Hidalgo • 1 ♀; Lagunilla ; [20.34° N, 99.02° W]; 14 Jun. 1951; H.E. Evans leg.; SEMC • 1 ♀; Zimapan ; [20.73° N, 99.38° W]; 11–14 Jun. 1951; P.D. Hurd leg.; EMEC • 1 ♂; same location as for preceding; 8 Jul. 1968; M.S. Wasbauer and J.E. Slansky leg.; UCDC . – Nuevo Leon • 2 ♀♀; 3 km SW of Galeana ; [24.81° N, 100.1° W]; 1550 m a.s.l.; 19 Mar. 1991; R. Brooks and R. Leschen leg.; SEMC • 1 ♀; 37 km SW of Linares ; [24.6° N, 99.8° W]; 310 m a.s.l.; 20 Mar. 1991; R. Brooks and R. Leschen leg.; SEMC • 6 ♀♀; Galeana , La Hediondilla; [25.0694° N, 100.7186° W]; 1890 m a.s.l.; 28 Jul. 2013; LRF and CVM leg.; LRF • 1 ♀; same location as for preceding; 2 Jun. 2013; LRF and CVM leg.; LRF • 1 ♀; same location as for preceding; 3 Apr. 2013; LRF and CVM leg.; LRF • 1 ♀; Galeana, Los Llanos ; [24.8089° N, 100.7119° W]; 1865 m a.s.l.; 27 Jul. 2013; LRF and CVM leg.; LRF • 5 ♀♀; same location as for preceding; 1 Jun. 2013; LRF and CVM leg.; LRF • 5 ♀♀; Galeana, Pablillo ; [24.6119° N, 100.0014° W]; 2085 m a.s.l.; 5 Apr. 2013; LRF and CVM leg.; LRF • 2 ♀♀; La Hediondilla, Galeana ; 25.0694° N, 100.7186° W; 1890 m a.s.l.; 29 Sep. 2013; LRF and CVM leg.; LRF • 1 ♀; Los Llanos, Galeana ; 24.8089° N, 100.7119° W; 1865 m a.s.l.; 29 Mar. 2014; LRF and CVM leg.; LRF • 1 ♀; Pablillo, Galeana ; 24.6119° N, 100.0014° W; 2083 m a.s.l.; 13 Apr. 2014; LRF and CVM leg.; LRF • 1 ♀; San Antonio de las Alazanas ; 25.2847° N, 100.6131° W; 2015 m a.s.l.; 1 Sep. 2013; LRF and CVM leg.; LRF . – San Luis Potosi • 1 ♀; El Huizache ; [22.92° N, 100.46° W]; 22 Aug. 1954; ex Larrea tridentata ; SEMC • 6 ♀♀; same location as for preceding; 22 Aug. 1954; SEMC . – Sonora • 1 ♀; 30 km E of Agua Prieta ; 31.3294° N, 109.2547° W; 24 Jul. 2001; R.L. Minckley leg.; RLM • 1 ♀; 30 km E of Agua Prieta ; 31.3322° N, 109.2703° W; 31 Mar. 2002; R.L. Minckley leg.; RLM • 4 ♀♀; Municipio de Agua Prieta, Rancho Las Anitas, Grassland Site 1; 31.31389° N, 109.31417° W; 21 May 2008; RLM • 1 ♀; Rancho San Bernardino, Bosque dry, Site 4; 31.30972° N, 109.27778° W; 8 May 2008; RLM • 1 ♀; Rancho San Bernardino, Bosque dry, Site 4, Transect 1; 31.30992° N, 109.27783° W; 19 May 2004; RLM • 1 ♀; Rancho San Bernardino, Bosque wet, Site 1, Transect 2; 31.33215° N, 109.27015° W; 3 Jul. 2002; RLM • 1 ♀; same location as for preceding; 13 Aug. 2002; RLM • 1 ♀; Rancho San Bernardino, field, Site 1, Transect 2; 31.32422° N, 109.26663° W; 11 Oct. 2002; RLM • 1 ♀; Rancho San Bernardino, Grassland, Site 1; 31.3187° N, 109.28023° W; 19 Aug. 2003; ex Kallstroemia californica ; RLM • 1 ♀; same location as for preceding; 19 Aug. 2003; RLM • 1 ♀; Rancho San Bernardino, Grassland, Site 6, Transect 1; 31.3112° N, 109.26625° W; 11 Aug. 2007; RLM • 1 ♀; Rancho San Bernardino, Grassland, Site 7; 31.30833° N, 109.27361° W; 12 Aug. 2006; RLM • 1 ♀; Rancho San Bernardino, Grassland, Site 7, Transect 3; 31.30865° N, 109.27308° W; 21 Jul. 2005; RLM • 1 ♀; Rancho San Bernardino, Little dam site, nr main houses; 31.32083° N, 109.27601° W; 16 Jul. 2005; ex Salsola kali subsp. tragus ; RLM • 1 ♀; Rancho San Bernardino, quarry, at road crossing; 31.32042° N, 109.2681° W; 2 Sep. 2003; RLM • 1 ♀; Rancho San Bernardino, quarry, gravel pit; 31.31964° N, 109.26926° W; 19 Aug. 2003; RLM • 1 ♀; Rancho San Bernardino, Rancho San Bernardino; 31.32907° N, 109.26553° W; 1 Aug. 2000; RLM • 1 ♀; Rancho San Bernardino, Riparian, Site 4; 31.32972° N, 109.25583° W; 20 Aug. 2003; RLM • 2 ♀♀; Rancho San Bernardino, Shrub 3; 31.32222° N, 109.29306° W; 7 Apr. 2006; RLM • 1 ♀; same location as for preceding; 23 Apr. 2008; RLM • 1 ♀; Rancho San Bernardino, Shrub, Site 4, Transect 2; 31.333° N, 109.24035° W; 6 Jul. 2002; RLM • 1 ♀; Sonora Rancho, San Bernardino; 31.33215° N, 109.27015° W; 5 Jul. 2001; R. Minckley leg.; RLM • 1 ♀; Sonora Rancho, San Bernardino; 31.3322° N, 109.2703° W; 1 Sep. 2001; R. Minckley leg.; RLM • 1 ♀; Sonora Rancho, San Bernardino; 31.3222° N, 109.2769° W; 11 Sep. 2001; R.L. Minckley leg.; RLM • 1 ♀; Sonora Rancho, San Bernardino; 31.3222° N, 109.2772° W; 22 Aug. 2001; R.L. Minckley leg.; RLM • 1 ♀; Sonora Rancho, San Bernardino; 31.3189° N, 109.2511° W; 31 Aug. 2001; R. Minckley leg.; RLM . – Tamaulipas • 1 ♀; 3 mi. S of San Fernando ; [24.8° N, 98.15° W]; 9 Jul. 1966; SEMC . – Zacatecas • 1 ♂; 45 mi. SW of Saltillo ; [24.7° N, 101.3° W]; 14 Sep. 1977; R.W. Brooks leg.; SEMC .

UNITED STATES – Arizona • 1 ♂; Apache Co., 33 km W of Zuni NM; [35.07° N, 109.21° W]; 14 Jun. 1999; L. Packer leg.; PCYU • 1 ♀; Cochise Co., 1 mi. S of Willcox; [32.24° N, 109.83° W]; 7 Aug. 1978; R.W. Brooks leg.; ex Wislizenia refracta ; SEMC • 1 ♀; Cochise Co., 4 mi. E of Willcox; [32.25° N, 109.76° W]; 17 Aug. 1994; J.G. Rozen and J.S.Ascher leg.; AMNH • 1 ♀; same location as for preceding; 11–18 Aug. 1987; K.V. Krombein leg.; USNM • 2 ♀♀; Cochise Co., 40 km S I-10 on Portal Rd; [31.87° N, 109.05° W]; 17 Apr. 1992; W.T. Wcislo leg.; ex Sisymbrium ; SEMC • 23 ♀♀; Cochise Co., Apache vicinity ; [31.69° N, 109.13° W]; 7 Jun. 1987; B.N. Danforth leg.; ex Lepidium ; SEMC • 2 ♂♂; Cochise Co., Chiricahua Mts , Desert Scrub Site ; [31.9° N, 109.3° W]; J.L. Neff and A.R. Moldenke leg.; CUIC • 1 ♀; Cochise Co., Douglas ; [31.34° N, 109.55° W]; 18–19 Jul. 1973; ex Larrea tridentata ; EMEC • 1 ♀; same location as for preceding; 18–19 Jul. 1973; P.D. Hurd, E.G. and J.M. Linsley, A.E. and M.M. Michelbacher leg.; ex Larrea tridentata ; EMEC • 1 ♀; same location as for preceding; 27 Jul. 1975; E.G. and J.M. Linsley leg.; ex Tamarix chinensis ; EMEC • 3 ♀♀; Cochise Co., Dry Lake , 4 mi. SE of Willcox, Sand dunes ; 32.2271° N, 109.7812° W; 27 Aug. 2013; T.L. Griswold leg.; BBSL • 1 ♀; Cochise Co., Portal Creek Lodge ; 31.9138° N, 109.1408° W; 1448 m a.s.l.; 5–6 Apr. 2016; C.M. Delphia leg.; MTEC • 1 ♀; Cochise Co., roadside, Out Side of Willcox; 32.2392° N, 109.7716° W; 10 May 2017; T. McMahon leg.; WRME • 1 ♀; Cochise Co., San Bernardino NWR ; 31.33627° N, 109.26752° W; 5 Jul. 2001; R. Minckley leg.; PCYU • 1 ♀; Cochise Co., San Bernardino NWR ; 31.3406° N, 109.2575° W; 31 Aug. 2001; R. Minckley leg.; RLM • 1 ♀; Cochise Co., San Bernardino NWR ; 31.3358° N, 109.2703° W; 13 Aug. 2001; R.L. Minckley leg.; RLM • 1 ♀; same location as for preceding; 31 Mar. 2002; R.L. Minckley leg.; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Bosque dry, Site 8; 31.33728° N, 109.23105° W; 4 Aug. 2003; RLM • 2 ♀♀; Cochise Co., San Bernardino NWR , Bosque dry, Site 8, Transect 2; 31.33735° N, 109.231° W; 18 Jul. 2005; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Bosque wet , Site 2; 31.33633° N, 109.2673° W; 22 Apr. 2007; RLM • 1 ♀; Cochise Co., San Bernardino NWR , field, Site 6 , Transect 4 ; 31.33444° N, 109.26806° W; 5 Jul. 2001; RLM • 1 ♀; Cochise Co., San Bernardino NWR , field, Site 8 , Transect 2 ; 31.3381° N, 109.2605° W; 19 Sep. 2001; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Grassland , Site 2 , Transect 3 ; 31.3358° N, 109.2702° W; 3 Jul. 2002; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Shrub , Site 5 , Transect 2 ; 31.34258° N, 109.27013° W; 30 Jul. 2004; RLM • 1 ♀; Cochise Co., Willcox ; [32.25° N, 109.83° W]; 18 Aug. 1958; P.D. Hurd leg.; ex Asclepias subverticillata ; EMEC • 1 ♀; same location as for preceding; 14 Aug. 1958; C.G. Moore leg.; UCDC • 1 ♀; Cochise Co., 6.4 km SE of Willcox on Blue Sky Rd; 32.2275° N, 109.7814° W; 1250 m a.s.l.; 25 Aug. 2001; R. Brooks leg.; SEMC • 1 ♀; same location as for preceding; 21 Aug. 2001; C. Michener leg.; ex Psorothamnus scoparius ; SEMC • 1 ♂; Pima Co., Silver Bell Bajada , IBP Desert Scrub Site ; [32.4° N, 111.5° W]; 17 Jul. 1974; J.L. Neff leg.; ex Heliotropium curassavicum ; CUIC • 1 ♀; Yavapai Co., 8 mi. NW of Wickenburg (Maricopa Co.); [34.05° N, 112.83° W]; 15 May 1992; J.G. and B.L. Rozen leg.; AMNH • 1 ♀; 20 mi. N of Flagstaff; [35.51° N, 111.55° W]; 2 Jul. 1952; C. Liang and W. LaBerge, R.H. and L.D. Beamer leg.; SEMC • 8 ♀♀, 1 ♂; 5 mi. E of Kingman; [35.19° N, 113.96° W]; 1 Jul. 1952; R.H. and L.D. Beamer, C. Weiner, A. Wolf, C. Liang and W. LaBerge leg.; SEMC • 1 ♀; 5 mi. W of Kingman; [35.19° N, 114.14° W]; 1 Jul. 1952; R.W. Brooks leg.; R.H. and L.D. Beamer, C. Weiner, A. Wolf, C. Liang and W. LaBerge leg.; SEMC • 3 ♀♀; 8 mi. SW of Willcox; [32.17° N, 109.93° W]; 12 Jul. 1952; R.H. and L.D. Beamer, Walley LaBerge and Cheng Liang leg.; SEMC • 1 ♀; Rustlers Pk. ; [31.905° N, 109.28° W]; 25 Jul. 1959; K.V. Krombein leg.; USNM. – California • 1 ♀; Riverside Co., Anza ; [33.56° N, 116.67° W]; 5 Jul. 1956; H.R. Moffitt leg.; UCDC • 1 ♀; Riverside Co., San Timoteo Canyon ; [33.98° N, 117.11° W]; 14 Aug. 1993; L. Packer and T. Pare leg.; PCYU • 1 ♀; San Bernardino Co., Cima Rd 6 mi. S I-15; 35.3667° N, 115.6° W; 1402 m a.s.l.; 1 Sep. 1994; R.R. Snelling leg.; ex Asclepias erosa ; LACM • 1 ♀; San Bernardino Co., Cima , 5.1 air mi. S of Sec. 31 T13N R14E; [35.17° N, 115.52° W]; 10 Jun. 1980; T. Griswold leg.; ex Eriogonum fasciculatum var. polifolium ; BBSL • 1 ♀; San Bernardino Co., Cotton Wash , Sec. 3 T9N R12 E; [34.9° N, 115.71° W]; 16 Jun. 1980; T. Griswold leg.; ex Acacia greggii ; BBSL • 1 ♀; San Bernardino Co., Granite Mts , T8N R13E Sec. 16; [34.78° N, 115.62° W]; 29 May 1981; Griswold leg.; BBSL • 1 ♀; San Bernardino Co., NE of Granite Mts, T9N R13E Sc33; [34.83° N, 115.62° W]; 1160 m a.s.l.; 21 Sep. 1993; T. Griswold leg.; ex Eriogonum deflexum ; BBSL • 1 ♀; San Bernardino Co., Valley Wells ; [35.79° N, 117.34° W]; 23 Apr. 1993; J.G. Rozen leg.; AMNH • 1 ♀; San Bernardino Co., Wild Horse Canyon , T11N R14E Sec. 1; [35.06° N, 115.44° W]; 23 Sep. 1993; T. Griswold leg.; ex Eriogonum deflexum ; BBSL • 1 ♀; San Bernardino Co., Winston Wash , Sec. 3 T10N R12E; [34.98° N, 115.71° W]; 588 m a.s.l.; 16 Apr. 1980; T. Griswold leg.; BBSL • 1 ♀; San Diego Co., Borrego Valley ; [33.26° N, 116.38° W]; 26 May 1954; F.X. Williams leg.; CAS • 5 ♀♀; Anza Borrego; 33.108° N, 116.4976° W; 20 Apr. 2013; Hung, Borchers and Nabor leg.; WRME • 8 ♀♀; same location as for preceding; 23 Mar. 2012; K.J. Hung leg.; WRME • 1 ♀; Anza-Borrego ; 33.05713° N, 116.4197° W; 13 Oct. 2014; K.J. Hung leg.; ex Isocoma acradenia ; WRME • 1 ♀; San Diego ; [32.7° N, 117.1° W]; 25 Jun. 2012; K. Hung leg.; WRME • 2 ♀♀; same location as for preceding; 3 May 2016; K.J. Hung leg.; WRME • 1 ♀; same location as for preceding; 31 Mar. 2016; Hung and Lee leg.; WRME • 1 ♀; same location as for preceding; 29 Jun. 2016; Hung and Lee leg.; WRME • 1 ♀; same location as for preceding; 28 Apr. 2015; Hung and Cen leg.; WRME • 1 ♀; Sunset Cove, Granites Res. Stn. Mojave Desert ; 34.77903° N, 115.66311° W; 12 Apr. 2017; J. Braun leg.; PCYU • 1 ♀; same location as for preceding; 15 Apr. 2017; J. Braun leg.; PCYU. – Colorado • 1 ♀; Boulder Co., Niwot Ridge Alpine Zone ; [40.06° N, 105.6° W]; 3 Jul. 1962; M.E. Smith leg.; UCMC • 1 ♀; Boulder Co., Open Space: Beech ; 40.0961° N, 105.2794° W; 7 Jul. 2004; Kearns and Oliveras leg.; PCYU • 1 ♂; El Paso Co., Colorado Springs ; 38.82539° N, 104.85979° W; 1886 m a.s.l.; 30 Jul. 2007; Gibbs and Sheffield leg.; PCYU • 1 ♀; Fremont Co., Coaldale ; [38.355° N, 105.785° W]; 2377 m a.s.l.; 4 Aug. 1957; C.D. Michener leg.; SEMC • 1 ♀; Huerfano Co., W of Walsenburg, Hwy 160; 37.6101° N, 104.81438° W; 1929 m a.s.l.; 4 Aug. 2007; Gibbs and Sheffield leg.; PCYU • 1 ♀; Jefferson Co.; 39.7476° N, 105.2453° W; 22 Aug. 2001; S.W. Droege leg.; PCYU • 1 ♀; Jefferson Co.; 39.731° N, 105.244° W; 21 Aug. 2001; S.W. Droege leg.; PCYU • 1 ♀; Lincoln Co., 17 mi. S of Limon; [39.02° N, 103.64° W]; 1585 m a.s.l.; 25 Jul. 1968; R.R. and R.A. Snelling leg.; ex Helianthus ; LACM • 18 ♀♀; Moffat Co., Artesia ; [40.24° N, 109.01° W]; 22 Jul. 1950; C.D. Michener leg.; ex Helianthus petiolaris ; SEMC • 3 ♀♀; Otero Co., Vogel Canyon ; [37.7° N, 103.5° W]; 12 Jun. 2008; W. Cranshaw leg.; CSUC • 1 ♀; Prowers Co., 1 mi. S of Granada; [38.05° N, 102.31° W]; 28 Aug. 1963; N. and B. Marston leg.; AMNH • 3 ♀♀; Prowers Co., Lamar , BB Road; [37.998° N, 102.7° W]; 13 Apr. 2002; H. Miller leg.; CSUC • 1 ♀; Weld Co., 10 mi. NE of Nunn Rd 45 and Rd 114; 40.81176° N, 104.6537° W; 1622 m a.s.l.; 11 Jun. 2001; V.L. Scott and S.Armstead leg.; ex Cleome serrulata ; WRME • 3 ♀♀; Weld Co., 10 mi. NE of Nunn Road 45 and Road 114; 40.81176° N, 104.6537° W; 1622 m a.s.l.; 11 Jun. 2001; Virginia Scott and S. Armstead leg.; ex Cleome serrulata ; UCMC • 1 ♂; Weld Co., Owl Creek, 12 mi. NE of Nunn; [40.86° N, 104.7° W]; 4 Aug. 1983; H.E. Evans leg.; CSUC • 1 ♀; same collection data as for preceding; CSUC • 1 ♂; Yuma Co., 4 mi. N. E. Idalia; [39.75° N, 102.24° W]; 10 Aug. 1964; J.G. and B.L. Rozen leg.; AMNH • 1 ♀; Boulder; [40.01° N, 105.27° W]; 23 May 1913; M.D. Ellis leg.; ex Capsella bursa-pastoris ; AMNH • 1 ♀; Julesburg ; 40.9833° N, 102.25° W; 1055 m a.s.l.; 7 Jun. 1920; AMNH • 2 ♀♀; Lamar; 38.1° N, 102.6167° W; 1097 m a.s.l.; 4–11 Jun. 1919; AMNH • 1 ♀; Lamar ; [38.09° N, 102.62° W]; 22 Jul. 1950; C.D. Michener leg.; ex Melilotus officinalis ; SEMC. – Idaho • 4 ♀♀; Canyon Co., Deer Flat NWR , DFO; 43.562° N, 116.66801° W; 11 Aug. 2011; Paul Backus leg.; FWSE • 3 ♀♀; same location as for preceding; 29 Jul. 2011; Addison Mohler leg.; FWSE • 5 ♀♀; same location as for preceding; 20 Apr. 2012; Addison Mohler leg.; FWSE • 4 ♀♀; same location as for preceding; 5 Jul. 2012; Sabrina Seidel leg.; FWSE • 4 ♀♀; Canyon Co., Deer Flat NWR , DFS; 43.55281° N, 116.64137° W; 5 Jul. 2012; Sabrina Seidel leg.; FWSE • 2 ♀♀; Glenn’s Ferry ; [42.95° N, 115.3° W]; 15 Sep. 1934; Bryant leg.; CAS • 1 ♀; Wendell, Pure Stand ; [42.77° N, 114.7° W]; 9 Jun. 1933; ex Sisymbrium / Salsola ; USNM. – Iowa • 1 ♀; Sioux City; [42.5° N, 96.4° W]; 17 Apr. 1937; C.N. Ainslie leg.; ex Salix ; AMNH. – Kansas • 17 ♀♀; Barber Co.; 37.015374° N, 98.371489° W; 27 Apr. 2013; S. Griffin leg.; WRME • 2 ♀♀; same location as for preceding; 28 Apr. 2013; S. Griffin leg.; WRME • 4 ♀♀; same location as for preceding; 29 Apr. 2013; S. Griffin leg.; WRME • 11 ♀♀; same location as for preceding; 30 Apr. 2013; S. Griffin leg.; WRME • 4 ♀♀; Barber Co.; 37.087305° N, 98.370901° W; 27 Apr. 2013; S. Griffin leg.; WRME • 1 ♀; same location as for preceding; 28 Apr. 2013; S. Griffin leg.; WRME • 2 ♀♀; same location as for preceding; 29 Apr. 2013; S. Griffin leg.; WRME • 4 ♀♀; same location as for preceding; 30 Apr. 2013; S. Griffin leg.; WRME • 1 ♀; Barber Co.; 37.034554° N, 98.366604° W; 27 Apr. 2013; S. Griffin leg.; WRME • 1 ♀; Reno Co., 5 mi. SE of Hutchinson; [38.01° N, 98° W]; 2 Sep. 1951; C.D. Michener and W.E. LaBerge leg.; ex Heterotheca ; UNSM. – Montana • 1 ♀; Choteau Co., Big Sandy , C1; 48.0443° N, 110.0695° W; 19 Jun. 2013; S. Adhikari leg.; MTEC • 3 ♀♀; same location as for preceding; 25 Jun. 2014; S. Adhikari leg.; MTEC • 1 ♀; Choteau Co., Big Sandy , C2; 48.0428° N, 110.0593° W; 25 Jun. 2014; S. Adhikari leg.; MTEC • 1 ♀; Choteau Co., Big Sandy , C3; 48.0383° N, 110.0593° W; 25 Jun. 2014; S. Adhikari leg.; MTEC • 1 ♀; Choteau Co., Big Sandy, O1; 48.0407° N, 110.0488° W; 8 Aug. 2014; S. Adhikari leg.; MTEC • 1 ♀; same location as for preceding; 5 Aug. 2015; S. Adhikari leg.; MTEC • 10 ♀♀; Choteau Co., Big Sandy , O2; 48.0363° N, 110.0383° W; 8 Aug. 2014; S. Adhikari leg.; MTEC • 8 ♀♀; same location as for preceding; 25 Jun. 2014; S. Adhikari leg.; MTEC • 1 ♀; same location as for preceding; 18 Jun. 2015; S. Adhikari leg.; MTEC • 1 ♀; same location as for preceding; 25 Jun. 2013; S. Adhikari leg.; MTEC • 1 ♀; Choteau Co., Big Sandy , O3; 48.0297° N, 110.0545° W; 8 Aug. 2014; S. Adhikari leg.; MTEC • 1 ♀; same location as for preceding; 25 Jun. 2014; S. Adhikari leg.; MTEC • 1 ♀; Gallatin Co., Gallatin Grown ; 45.7713° N, 111.2953° W; 9 Jul. 2015; C.M. Delphia and J.L. Monte leg.; MTEC • 2 ♀♀; same location as for preceding; 13 Aug. 2015; C.M. Delphia and J.L. Monte leg.; MTEC • 1 ♀; same location as for preceding; 24 Jul. 2015; C.M. Delphia and J.L. Monte leg.; MTEC • 1 ♀; same location as for preceding; 25 Jul. 2013; C.M. Delphia leg.; MTEC • 1 ♀; same location as for preceding; 8 Aug. 2013; C.M. Delphia leg.; MTEC • 1 ♀; same location as for preceding; 22 Aug. 2013; C.M. Delphia leg.; MTEC • 1 ♀; same location as for preceding; 19 Jul. 2013; C.M. Delphia leg.; MTEC • 1 ♀; Gallatin Co., MSU BART Farm ; 45.6675° N, 111.0663° W; 29 Jul. 2010; C.M. Delphia leg.; ex Campanula rotundifolia ; MTEC • 1 ♀; Gallatin Co., Rocky Creek Farm , 34297 Frontage Road; [45.666° N, 110.954° W]; 8 Aug. 2011; C.M. Delphia leg.; MTEC • 1 ♀; Rosebud Co., Four Wing Saltbush , 3 mi. S of Colstrip; [45.84° N, 106.62° W]; 10 Aug. 1983; MTEC • 1 ♀; same location as for preceding; 13 Sep. 1983; MTEC • 1 ♀; Rosebud Co., Rosebud Rest Stop ; [46.265° N, 106.3° W]; 22 Jun. 1987; T.A. Ebert leg.; CSUC. – Nebraska • 1 ♀; Lincoln Co., 8–14 mi. NNW of North Platte; [41.3° N, 100.9° W]; 8 Aug. 1971; J.G., B.L. and K.C. Rozen leg.; WRME • 4 ♀♀; McPherson Co., Sandhills Ag Lab ; [41.62° N, 100.83° W]; 28 Jun.–10 Aug. 1972; UNSM • 2 ♀♀; same location as for preceding; 15 Jul. 1972; UNSM • 4 ♀♀; same location as for preceding; 20 Jul. 1972; UNSM • 1 ♂; 10 mi. SW of Imperial; [40.41° N, 101.78° W]; 5 Sep. 1961; Crew leg.; ex misc. vegetation; UNSM • 1 ♀; 3 mi. E of Thedford; [41.98° N, 100.52° W]; 19 Aug. 1960; W.E. LaBerge leg.; ex Euphorbia marginata ; UNSM • 1 ♂; 3 mi. S of Hyannis; [41.96° N, 101.76° W]; 12 Sep. 1961; Crew leg.; ex misc. vegetation; UNSM. – Nevada • 1 ♀; Clark Co., 0.3 mi. SE of Corn Creek Spr.; 36.4365° N, 115.3525° W; 901 m a.s.l.; 24 Aug. 2004; S. Higbee and E. Ahlstrom leg.; BBSL • 1 ♀; Clark Co., 1.4 mi. WSW of Bonanza Hill; 35.7623° N, 115.5796° W; 792 m a.s.l.; 4 May 2004; Higbee, Ahlstrom, Saul leg.; BBSL • 1 ♀; Clark Co., 2.5 mi. SE of Rainbow Mtn; 36.0896° N, 115.4493° W; 1138 m a.s.l.; 9 Sep. 2004; S. Higbee and T. Griswold leg.; BBSL • 1 ♀; Clark Co., 0.69 mi. N of Beehive Rock; 36.4196° N, 114.5509° W; 23 Aug. 2005; E. North leg.; ex Croton wigginsii ; WRME • 1 ♀; Clark Co., Peek-A-Boo Canyon ; 36.5326° N, 115.1117° W; 1760 m a.s.l.; 16 Aug. 2004; S. Higbee leg.; ex Eriogonum cernuum ; BBSL • 1 ♀; Clark Co., 2.85 mi. SE of Rainbow Mtn; 36.0804° N, 115.4587° W; 24 Jun. 2005; S. Higbee leg.; PCYU • 1 ♀; Clark Co., Wheeler Wash ; 36.2748° N, 115.8425° W; 1591 m a.s.l.; 9 Jun. 2004; E. Ahlstrom leg.; ex Baileya multiradiata ; BBSL • 3 ♀♀; Elko Co., Elko ; [40.83° N, 115.76° W]; 7 Jun. 1982; R.W. Brooks leg.; ex Sphaeralcea ambigua ; SEMC. – New Mexico • 2 ♀♀; Bernalillo Co., Cibola NF , Sandia Crest , Kiwanis Meadow ; 35.2058° N, 106.4423° W; 3057 m a.s.l.; 21 Jun. 2017; J.A. Fowler leg.; UCRC • 1 ♀; Bernalillo Co., Cibola NF , Sandia Crest , Kiwanis Meadow ; 35.2059° N, 106.4431° W; 3057 m a.s.l.; 21 Jun. 2017; J.A. Fowler leg.; UCRC • 1 ♀; Bernalillo Co., Cibola NF , Sandia Crest , Kiwanis Meadow ; 35.2052° N, 106.4424° W; 3057 m a.s.l.; 21 Jun. 2017; J.A. Fowler leg.; UCRC • 2 ♀♀; same location as for preceding; 22 Jun. 2017; J.A. Fowler leg.; UCRC • 1 ♀; Bernalillo Co., Cibola NF , Sandia Crest , Kiwanis Meadow ; 35.2059° N, 106.4431° W; 3057 m a.s.l.; 22 Jun. 2017; J.A. Fowler leg.; UCRC • 1 ♀, 1 ♂; Chaves Co., 5 km S of Roswell, Hwy 285; 33.36177° N, 104.51238° W; 1108 m a.s.l.; 10 Aug. 2007; Gibbs and Sheffield leg.; PCYU • 1 ♂; Chaves Co., Bottomless Lakes , 10 mi. E of Roswell; [33.34° N, 104.34° W]; 20 May 1969; Brothers, Krueger and Michener leg.; ex Tamarix gallica ; SEMC • 1 ♀; Colfax Co., 7.6 mi. NW of Kiowa; 36.73724° N, 104.22277° W; 2195 m a.s.l.; 18 Sep. 1999; A.L. Hicks and Virginia Scott leg.; ex Schizachyrium scoparium ; UCMC • 2 ♀♀; Colfax Co., Hwy 64; 36.63692° N, 104.67754° W; 1932 m a.s.l.; 6 Aug. 2007; Gibbs and Sheffield leg.; PCYU • 1 ♀; DeBaca Co., 10 mi. S of Ft Sumner; [34.32° N, 104.25° W]; 19 May 1969; Brothers, Krueger and Michener leg.; ex Sphaeralcea ; SEMC • 1 ♀; same location as for preceding; 19 May 1969; Brothers, Krueger and Michener leg.; ex Gaillardia ; SEMC • 1 ♂; Dona Ana Co., 16 mi. N of Las Cruces; [32.55° N, 106.77° W]; 18 Aug. 1993; J.W. Van Zee leg.; ex Gutierrezia ; NMSU • 1 ♂; same location as for preceding; 1 Sep. 1993; J.W. Van Zee leg.; NMSU • 1 ♀; Dona Ana Co., jct. West Afton Rd and Black Mt. Rd; 32.0908° N, 106.7794° W; 1272 m a.s.l.; 14 Sep. 2008; G.R. Ballmer leg.; ex Gutierrezia ; UCRC • 1 ♀; Dona Ana Co., Jornada Range 26 km N of Las Cruces, stressor Site 8 km W of USDA HQ, Plot 17; [32.61° N, 106.83° W]; 1200 m a.s.l.; 4 Sep. 1996; G. Forbes leg.; NMSU • 1 ♀; Dona Ana Co., Jornada Range 26 km N of Las Cruces, stressor Site 8 km W of USDA HQ, Plot 7; [32.61° N, 106.83° W]; 1200 m a.s.l.; 11 Aug. 1997; G. Forbes leg.; NMSU • 1 ♀; Dona Ana Co., Las Cruces ; [32.32° N, 106.76° W]; 15 Aug. 1967; R.M. Bohart leg.; UCDC • 2 ♀♀, 1 ♂; Eddy Co., 22 mi. SW of Carlsbad; [32.2° N, 104.5° W]; 21 May 1969; ex Lepidium integrifolium ; SEMC • 1 ♀, 1 ♂; same location as for preceding; 21 May 1969; Brothers, Krueger and Michener leg.; ex Chamaesaracha ; SEMC • 1 ♀; Eddy Co., 3 mi. S of Artesia; [32.8° N, 104.4° W]; 20 May 1969; Brothers, Krueger and Michener leg.; ex Chamaesaracha coniodes ; SEMC • 2 ♀♀; Eddy Co., Carlsbad ; [32.42° N, 104.23° W]; 20 May 1969; Brothers, Krueger and Michener leg.; ex Lepidium integrifolium ; SEMC • 3 ♀♀; Eddy Co.; 32.1° N, 104.5° W; 1109 m a.s.l.; 10 Jun. 2010; J.D. Herndon and A. Druk leg.; BBSL • 1 ♀; Eddy Co.; 32.1° N, 104.6° W; 1551 m a.s.l.; 9 Jun. 2010; J.D. Herndon and A. Druk leg.; BBSL • 1 ♀; same location as for preceding; 1 Sep. 2010; R. Krauss and J.D. Herndon leg.; BBSL • 1 ♀; Eddy Co.; 32.1638° N, 104.4443° W; 1152 m a.s.l.; 31 Aug. 2010; J.D. Herndon and R. Krauss leg.; BBSL • 2 ♀♀; Eddy Co.; 32.1768° N, 104.4407° W; 30 Aug. 2010; R. Krauss leg.; BBSL • 1 ♀; Eddy Co.; 32.1117° N, 104.5631° W; 1324 m a.s.l.; 12 Jul. 2010; J.D. Herndon and A. Druk leg.; BBSL • 1 ♀; Eddy Co.; 32.1856° N, 104.4326° W; 7 Apr. 2010; H. Ikerd and T.L. Griswold leg.; BBSL • 2 ♀♀; Hidalgo Co., 1.6 km S of Rodeo; [31.82° N, 109.03° W]; 1 Aug. 1987; B.N. Danforth leg.; ex Eriogonum ; SEMC • 2 ♀♀; Hidalgo Co., 12.3 mi. N of Rodeo; [32.01° N, 109.03° W]; Aug. 1986; R.J. McGinley and B.M. Norden leg.; USNM • 1 ♀; Hidalgo Co., 13 mi. S of Animas; 31.7656° N, 108.8069° W; 1219 m a.s.l.; 19 Aug. 2001; SEMC • 2 ♂♂; Hidalgo Co., 20 mi. S of Animas; [31.66° N, 108.81° W]; 23 Aug. 2008; J.S. Ascher, J.G. Rozen and M.A. Rozen leg.; AMNH • 1 ♂; Hidalgo Co., 23.5 mi. S of Animas; 31.61838° N, 108.86737° W; 23 Aug. 2007; J.J. Gibbs leg.; PCYU • 1 ♂; Hidalgo Co., 24.5 mi. S of Animas; [31.59° N, 108.81° W]; 23 Aug. 2007; J.J. Gibbs leg.; PCYU • 12 ♀♀, 1 ♂; Hidalgo Co., 9 mi. N of Cotton City; [32.21° N, 108.88° W]; 11 Aug. 1978; R.W. Brooks leg.; ex Wislizenia refracta ; SEMC • 2 ♀♀; Hidalgo Co., Animas, 8.8 km E of junc. Sock N Shoe Rd.; 31.9572° N, 108.7144° W; 1390 m a.s.l.; 15 Aug. 2000; R. Brooks leg.; ex Fallugia paradoxa ; SEMC • 3 ♂♂; Lea Co., 30 mi. W of Eunice; 32.5° N, 103.7° W; 17 May 2015; P. Rude leg.; EMEC • 4 ♀♀; Lincoln Co., N Carrizozo on Rte 54; [33.8° N, 105.84° W]; 15 May 1987; B.N. Danforth leg.; ex Physaria ; SEMC • 1 ♀; Otero Co., Lincoln NF , 2 mi. E of Sunspot, meadow; 32.7886° N, 105.786° W; 11 Jul. 2017; J.A. Fowler leg.; UCRC • 1 ♀; Roosevelt Co., nr Boone Draw , 14 mi. SW of Portales; [34.05° N, 103.52° W]; 26 May 1972; Saul Frommer and N. Jorgensen leg.; UCRC • 1 ♀; Santa Fe Co., Santa Fe NF, Jack’s Creek Campground , meadow ; 35.8555° N, 105.6398° W; 3042 m a.s.l.; 24 Jul. 2017; J.A. Fowler leg.; UCRC • 1 ♀; Socorro Co., Bosque del Apache ; [33.8° N, 106.89° W]; 1375 m a.s.l.; 15 Jul. 1991; Ilan Yarom leg.; SEMC • 3 ♀♀; Socorro Co., Cibola NF, South Baldy meadow ; 33.9901° N, 107.1841° W; 3178 m a.s.l.; 16 Jun. 2017; K.C. Tom leg.; UCRC • 2 ♀♀; Socorro Co., Cibola NF , South Baldy meadow ; 33.9919° N, 107.1836° W; 3178 m a.s.l.; 16 Jun. 2017; M P. Argueta leg.; UCRC • 2 ♀♀; Socorro Co., Cibola NF , South Baldy meadow ; 33.9911° N, 107.1837° W; 3178 m a.s.l.; 16 Jun. 2017; J.A. Fowler leg.; UCRC • 1 ♀; Socorro Co., Sevilleta NWR; [34.3° N, 106.8° W]; 18 Apr.–2 May 2006; K. Wetherill leg.; PCYU • 1 ♀; same location as for preceding; 17–31 May 2005; K. Wetherill leg.; PCYU • 1 ♀; same location as for preceding; 5–19 Oct. 2004; K. Wetherill leg.; PCYU • 1 ♀; Torrance Co., 5 mi. NW of Cedarvale; [34.42° N, 105.76° W]; 6 Aug. 1966; C.R. Kovacic leg.; UCDC • 1 ♀; Valencia Co., ASC Los Lunas ; [34.81° N, 106.73° W]; 25 Apr. 1988; C.R. Ward leg.; NMSU • 1 ♀; Valencia Co., Los Lunas; [34.81° N, 106.73° W]; 10 May 1988; C.R. Ward leg.; NMSU • 1 ♀; 25 mi. W of Tularosa; [33.07° N, 106.45° W]; 1 Jul. 1940; L.J. Lipovsky leg.; SEMC • 4 ♀♀; same location as for preceding; 1 Jul. 1940; D.E. Hardy leg.; SEMC • 1 ♀; Alamogordo ; [32.9° N, 105.96° W]; 26 Jun. 1940; D.E. Hardy leg.; SEMC • 1 ♀; same location as for preceding; 26 Jun. 1940; L.J. Lipovsky leg.; SEMC • 1 ♀; Bernalillo ; [35.3° N, 106.55° W]; 23 Jul. 1950; L.D. Beamer leg.; SEMC • 1 ♀; Carrizozo; [33.64° N, 105.88° W]; 9 Jun. 1950; L.D. Beamer leg.; ex Mertensia franciscana ; SEMC • 1 ♀; same location as for preceding; 9 Jun. 1950; L.D. Beamer leg.; ex Asclepias verticillata ; SEMC • 2 ♀♀; same location as for preceding; 9 Jun. 1950; L.D. Beamer leg.; SEMC • 1 ♀; Cloudcroft ; [32.96° N, 105.74° W]; 14 Jul. 1936; J.D. Beamer leg.; SEMC • 1 ♀; Cuervo ; [35.03° N, 104.41° W]; 23 Jun. 1940; E.E. Kenaga leg.; SEMC • 1 ♀; Las Cruces; [32.32° N, 106.76° W]; 12 Jun. 1950; R.H. Beamer leg.; SEMC • 1 ♀; Organ ; [32.43° N, 106.6° W]; 3 Jul. 1940; D.E. Hardy leg.; SEMC • 1 ♂; Eddy Co.; 32.1095° N, 104.4716° W; 1115 m a.s.l.; 19 Jul. 2011; N. Klass leg.; ex Chilopsis linearis ; BBSL • 1 ♂; Eddy Co.; 32.1669° N, 104.5028° W; 1373 m a.s.l.; 27 Jun. 2011; J.D. Herndon and N. Klass leg.; BBSL. – North Dakota • 1 ♀; Linton ; [46.27° N, 100.23° W]; 13 May 1940; D.G. Denning leg.; UMSP • 1 ♀; Barnes Co., 6.6 km NW of Sandborn; 46.9794° N, 98.2916° W; 14 Jun. 2011; E. Evans leg.; UMSP. – Oklahoma • 1 ♀; Cleveland Co.; [36.31° N, 96.46° W]; 6 Jul. 1981; Pat Terry leg.; ex Aphanostephus skirrhobasis ; UNSM • 2 ♀♀; same location as for preceding; 7 Jul. 1981; Pat Terry leg.; ex Aphanostephus skirrhobasis ; UNSM • 3 ♀♀; Garfield Co., Enid; [36.4° N, 97.88° W]; 14 Apr. 1961; Rozen and Schrammel leg.; AMNH. – South Dakota • 1 ♀; Pennington Co., Badlands National Park : North Unit ; 43.8421° N, 102.2024° W; 812 m a.s.l.; 27 Jun. 2002; Laura Schoessler leg.; UMSP • 1 ♂; Cedar Pass, Bad Lands ; [43.757° N, 101.936° W]; 14 Aug. 1940; H.E. Milliron leg.; UMSP. – Texas • 1 ♂; Bailey Co., Muleshoe Wildlife Refuge , Grassland 1; 33.963° N, 102.7282° W; 14 May 2013; S.J. Galdek leg.; BBSL • 1 ♂; same location as for preceding; 29 Apr. 2013; S.J. Galdek leg.; BBSL • 1 ♂; Bailey Co., Muleshoe Wildlife Refuge , Prairie Dog 2; 33.9302° N, 102.7584° W; 10 Jun. 2013; S.J. Galdek leg.; BBSL • 1 ♂; Bailey Co., Muleshoe Wildlife Refuge , Prairie Dog 3; 33.963° N, 102.7749° W; 30 Apr. 2013; S.J. Galdek leg.; BBSL • 1 ♀; Brewster Co., Big Bend Nat’l. Park , Cattail Falls ; [29.273° N, 103.336° W]; 9 Jun. 1999; S. Droege leg.; BBSL • 1 ♀; Brewster Co., Calamity Creek Rd., 25 km S of Alpine; [30.14° N, 103.63° W]; 29 Apr. 2003; Packer and Fraser leg.; PCYU • 2 ♀♀; Brewster Co., Marathon; [30.21° N, 103.24° W]; 30 May 1993; R.L. Minckley and K. Toal leg.; ex Sphaeralcea leptophylla ; SEMC • 1 ♀; Colorado Co.; [29.6° N, 96.5° W]; 30 Mar. 1922; Mrs. Grace Wiley leg.; SEMC • 1 ♀; same location as for preceding; 7Apr. 1922; Mrs Grace Wiley leg.; SEMC • 1 ♀; Culberson Co., Hurds Draw Rd and Hwy 10 intersection; [31.07° N, 104.28° W]; 30 May 1999; L. Packer leg.; PCYU • 5 ♀♀; Denton Co.; 33.2043° N, 97.0816° W; 27 Apr. 2002; H.W. Ikerd leg.; PCYU • 1 ♀; Dimmit Co., Chaparral Wildlife Management Area ; [28.33° N, 99.41° W]; 20 Aug. 1994; J.L. Neff and A. Hook leg.; ex Amphiachyris dracunculoides ; CTMI • 2 ♀♀; Ellis Co., Waxahachie ; [32.39° N, 96.85° W]; 26 Jun. 1988; J.G. Rozen leg.; AMNH • 1 ♀; Hall Co., 6 mi. SE of Turkey; [34.33° N, 100.97° W]; 28 May 1970; C.W. O’Brien leg.; LACM • 1 ♀; Hall Co., 16 mi. S of Memphis; [34.49° N, 100.54° W]; 22 Jun. 1959; R.R. Snelling and M.D. Snelling leg.; ex Tamarix gallica ; LACM • 1 ♀; Hidalgo Co., Bentsen Rio Grande SP ; 26.186° N, 98.374° W; 16 Mar. 2007; J.L. Neff leg.; ex Nama sp.; CTMI • 1 ♀; Hidalgo Co., Bentsen-Rio Grande State Park ; [26.175° N, 98.385° W]; 29 Mar. 1991; J L. Neff leg.; CTMI • 1 ♀; Hudspeth Co., Fort Hancock ; 31.3028° N, 105.8395° W; 17 Aug. 2000; J.L. Neff leg.; CTMI • 1 ♀, 3 ♂♂; Jeff Davis Co., 0.5 mi. S of Kent; [31.06° N, 104.22° W]; 29 May 1999; L. Packer leg.; PCYU • 1 ♀; Jeff Davis Co., 20 km S of Kent; [30.89° N, 104.22° W]; 30 Apr. 2003; Packer and Fraser leg.; PCYU • 3 ♀♀; Jeff Davis Co., 9 mi. S of Kent; [30.94° N, 104.22° W]; 29 May 1999; L. Packer leg.; PCYU • 1 ♀; Jeff Davis Co., 9 mi. SE of Fort Davis; [30.5° N, 103.79° W]; 23 Aug. 1967; R.C. Gardner and C.R. Kovacic leg.; UCDC • 2 ♀♀; Jeff Davis Co., Davis Mts TNC Res .; 30.708° N, 104.099° W; 1800 m a.s.l.; 16 May 2005; J L. Neff and A. Hook leg.; CTMI • 1 ♀; Kimble Co., 19 mi. NNW of Harper; 30.483° N, 99.302° W; 28 May 2006; J.L. Neff and A. Hook leg.; ex Thelesperma filifolium ; CTMI • 1 ♀; Kimble Co., RM 385 and Salt Branch Rd; 30.478° N, 99.402° W; 2 Jul. 2005; A.W. Hook and J.L. Neff leg.; CTMI • 1 ♀; Kimble Co., Salt Branch Rd.; 30.478° N, 99.402° W; 6 Jun. 2004; A.W. Hook and J.L. Neff leg.; CTMI • 1 ♀; La Salle Co., Chaparral Wildlife Management Area ; [28.33° N, 99.39° W]; 15 May 2003; L. Packer leg.; PCYU • 3 ♀♀, 1 ♂; same location as for preceding; 15–16 May 2003; L. Packer leg.; PCYU • 1 ♂; La Salle Co.; 15 May 2003; L. Packer leg.; PCYU • 1 ♀; Mason Co., Salt Branch X River Rds ; 30.511° N, 99.31° W; 28 May 2006; J.L. Neff and A. Hook leg.; ex Erigeron modestus ; CTMI • 1 ♂; Randall Co., Buffalo Lake Wildlife Refuge , Grassland 2 ; 34.8892° N, 102.1084° W; 7 May 2013; S.J. Galdek leg.; BBSL • 1 ♂; Randall Co., Buffalo Lake Wildlife Refuge , Prairie Dog 1 ; 34.9028° N, 102.0989° W; 3 Jun. 2013; S.J. Galdek leg.; BBSL • 1 ♂; Randall Co., Buffalo Lake Wildlife Refuge, Prairie Dog 2 ; 34.8896° N, 102.1084° W; 4 Jun. 2013; S.J. Galdek leg.; BBSL • 1 ♂; same location as for preceding; 3 Jun. 2013; S.J. Galdek leg.; BBSL • 1 ♂; Randall Co., Buffalo Lake Wildlife Refuge , Prairie Dog 3 ; 34.8713° N, 102.1492° W; 6 Jun. 2013; S.J. Galdek leg.; BBSL • 2 ♀♀; Reeves Co., 3 mi. NW of Pecos; 31.453° N, 103.508° W; 6 Jul. 2007; J.L. Neff and A. Hook leg.; ex Lepidium alyssoides ; CTMI • 1 ♀; San Patricio Co., Welder Refuge ; 28.1122° N, 97.4167° W; Mar. 2004; Sty Reed leg.; PCYU • 1 ♀; 10 mi. N of Mason; [30.89° N, 99.23° W]; 13 Apr. 1952; Michener, Beamers, Wille and LaBerge leg.; ex Physaria ; LACM • 1 ♀; Lubbock ; [33.58° N, 101.86° W]; 19 Feb. 1967; V. Barber leg.; LACM. – Utah • 1 ♀; Emery Co., 9 km N of Goblin Valley St. Pk.; [38.65° N, 110.66° W]; 1360 m a.s.l.; 14 Aug. 1992; D. Yanega leg.; SEMC • 1 ♀; same location as for preceding; 14 Aug. 1992; R. Minckley leg.; ex Lathrocasis tenerrima ?; SEMC • 11 ♀♀; Emery Co., Green River ; 38.9885° N, 110.162° W; 9 May 2000; F.D. Parker leg.; BBSL • 1 ♀; Emery Co., San Rafael Desert , 1 mi. E of Flat Top Pass; 38.5417° N, 110.465° W; 23 May 2000; F.D. Parker leg.; BBSL • 1 ♀; Emery Co., San Rafael Desert, 3 mi. E of Flat Top Pass; 38.51° N, 110.4255° W; 24 May 2000; F.D. Parker leg.; BBSL • 1 ♀; Garfield Co., 2.3 mi. S of Pioneer Mesa; 37.7622° N, 111.1293° W; 20 May 2002; C. Davidson leg.; ex Fallugia paradoxa ; BBSL • 1 ♀; Grand Co., 8 mi. SE of Emery; [38.84° N, 111.14° W]; 19 May 1980; D.B. Wahl leg.; SEMC • 1 ♀; same location as for preceding; 22 May 1979; D.B. Wahl leg.; SEMC • 1 ♀; Kane Co., 1.32 mi. SSW of Cougar Knoll; 37.2911° N, 111.1414° W; 7 Jul. 2000; O. Messinger leg.; ex Potentilla gracilis ; BBSL • 1 ♀; Kane Co., 1.33 mi. SSW of Cougar Knoll; 37.2931° N, 111.1505° W; 7 Jul. 2000; C. Davidson leg.; ex yellow composite; PCYU • 1 ♀; Kane Co., 2.3 mi. NNW of White Point; 37.5452° N, 111.322° W; 17 Sep. 2001; K. Moredock leg.; ex Ericameria nauseosa var. nauseosa ; PCYU • 1 ♀; Kane Co., 2.7 mi. SE of Left Hand Collet Canyon; 37.5085° N, 111.4069°W; 13 May 2002; K. Fordney leg.; ex Amelanchier utahensis ; BBSL • 1 ♀; Kane Co., Tibbet Canyon ; 37.1606° N, 111.5392° W; 16 May 2002; S. Higbee leg.; ex Sphaeralcea grossulariifolia ; BBSL • 2 ♀♀; Millard Co., Kanosh ; [38.8° N, 112.44° W]; 24 May 1963; G.F. Knowlton leg.; SEMC • 1 ♀; Millard Co., Oak Creek Canyon ; [39.35° N, 112.26° W]; 27 May 1958; G.F. Knowlton leg.; ex Salix sp.; SEMC • 1 ♀; Washington Co., 0.72 mi. E of Firepit Knoll; 37.3494° N, 113.0912° W; 19 Jun. 2006; B. Hays and F. Nicklen leg.; PCYU • 1 ♀; Washington Co., 1.1 mi. N of Rockville; 37.1773° N, 113.0368° W; 11 May 2006; B. Hays leg.; PCYU • 1 ♀; Cedar Breaks ; [37.63° N, 112.85° W]; 9 Mar. 1992; L. Packer leg.; PCYU • 1 ♀; Goshen; [39.95° N, 111.9° W]; 12Aug. 1949; Geo. E. Bohart leg.; BBSL • 1 ♀; Kane Spgs., 15 mi. SE of Moab; [38.392° N, 109.452° W]; 9 Aug. 1957; C.D. Michener leg.; ex Cleome lutea ; SEMC • 1 ♀; Roosevelt ; [40.3° N, 109.99° W]; 29 Jun. 1954; G.F. Knowlton leg.; SEMC • 1 ♀; Tremonton ; [41.71° N, 112.17° W]; 5 Jul. 1931; J.O. Nottingham leg.; SEMC. – Wyoming • 1 ♀; Big Horn Co., 4 mi. southwest Lane 13, Lovell Local Training Area ; [44.77° N, 108.3° W]; 1 Jun. 2004; B. Kondratieff and J. Owens leg.; CSUC • 1 ♀; Hot Springs Co., 10 mi. N of Thermopolis; [43.79° N, 108.21° W]; 12 Aug. 1975; C.D. Michener leg.; ex Cleome lutea ; SEMC • 1 ♀; Natrona Co., 25 mi. SW of Casper; [42.59° N, 106.67° W]; 12 Aug. 1972; J.G. Rozen and R. McGinley leg.; AMNH • 1 ♀; Natrona Co., 9 mi. E. Casper; [42.85° N, 106.14° W]; 12 Aug. 1962; J.G. and B.L. Rozen leg.; AMNH • 4 ♀♀; Platte Co., Warm Spring , Camp Guernsey ; [42.26° N, 104.76° W]; 1372 m a.s.l.; 6 Jun. 2000; P.M. Pineda and J. Schmidt leg.; CSUC • 1 ♀; Sheridan Co., 5 mi. N of Sheridan; [44.87° N, 106.96° W]; 6 Aug. 1963; G. Bohart and P. Torchio leg.; ex Solidago ; BBSL • 1 ♀; Grand Teton Nat. Pk. ; [43.8° N, 110.7° W]; Jul. 1937; BBSL.

Description

See Gibbs (2010) for a complete redescription and figures.

Range

Alberta east to North Dakota, south to Baja California Sur and San Luis Potosi ( Fig. 42 ).

Floral records

AMARANTHACEAE Juss. Salsola L. S. kali L. S. k. subsp. tragus (L.) Čelak. • APOCYNACEAE Juss. Asclepias L. A. erosa Torr. • A. subverticillata (A.Gray) Vail • A. verticillata L. • ASTERACEAE Giseke : Amphiachyris : A. dracunculoides (DC.) Nutt. • Aphanostephus : A. skirrhobasis (DC.) Trel. ex Coville & Branner • Bahia : B. absinthifolia Benth. • Baileya : B. multiradiata Harv. &A.Gray ex Torr. • Ericameria : E. nauseosa (Pall. ex Pursh) G.L. Nesom & Baird : E. n. var. nauseosa (Pall. ex Pursh) G.L. Nesom & Baird • Erigeron : E. modestus A.Gray • Gaillardia • Gutierrezia • Helianthus : H. petiolaris Nutt. • Heterotheca • Isocoma : I. acradenia (Greene) Greene • Solidago • Thelesperma : T. filifolium (Hook.) A.Gray • Varilla : V. mexicana A.Gray • BIGONIACEAE Juss. Chilopsis D. Don : C. linearis (Cav.) Sweet • BORAGINACEAE Juss. Heliotropium L. H. curassavicum L. • Mertensia Roth : M. franciscana A. Heller • Nama L. • BRASSICACEAE Burnett : Brassica L. B. nigra (L.) K. Koch • Capsella Medik. C. bursa-pastoris (L.) Medik. • Lepidium L. L. alyssoides A.Gray • L. integrifolium Nutt. • Physaria (Nutt. ex Torr. & A.Gray) A.Gray • Sisymbrium L. • CAMPANULACEAE Juss. Campanula L. C. rotundifolia L. • CLEOMACEAE Bercht. & J. Presl : Cleome L. C. lutea Hook. • C. serrulata Pursh • Wislizenia Engelm. W. refracta Engelm. • EUPHORBIACEAE Juss. Croton L. C. wigginsii L.C. Wheeler • Euphorbia L. E. marginata Pursh • FABACEAE Juss. Acacia : A. greggii A.Gray • Dalea : D. eriophylla S. Watson • Melilotus : M. officinalis (L.) Pall. • Psorothamnus : P. scoparius (A.Gray) Rydb. • MALVACEAE Juss. Sphaeralcea A. St. -Hil. S. ambigua A.Gray • S. grossulariifolia (Hook. & Arn.) Rydb. • S. leptophylla (A.Gray) Rydb. • POACEAE Barnhart : Schizachyrium Nees : S. scoparium (Michx.) Nash • POLYGONACEAE Juss. Eriogonum Michx. E. cernuum Nutt. • E. deflexum Torr. • E. fasciculatum Benth. E. f. var. polifolium (Benth.) Torr. & A.Gray • ROSACEAE Juss. Amelanchier Medik. A. utahensis Koehne • Fallugia Endl. F. paradoxa (D. Don) Endl. ex Torr. • Potentilla L. P. gracilis Douglas ex Hook. • SALICACEAE Mirb. Salix L. • SOLANACEAE Adans. Chamaesaracha Franch. & Sav. ex A.Gray : C. coniodes (Dunal) Britton • TAMARICACEAE Link : Tamarix L. T. chinensis Lour. • T. gallica L. • ZYGOPHYLLACEAE R.Br. Kallstroemia Scop. K. californica (S. Watson) Vail • Larrea Cav. L. tridentata (Sessé & Moc. ex DC.) Coville.

DNA barcodes

Twnety-four sequences available (BOLD process IDs: DLII021-06, DLII052-06, DLII785-07, DLII801-07, DLII851-07, DLII852-07, DLII874-07, DLII912-07, DLII913-07, DLII983-07, DLII999- 07, DLII1032-07, DLII1079-07, DLII1083-07, DLII1125-07, DLII1127-07, DLII1266-08, DLII1273- 08, DLII1472-08, DLIII103-18, DIAL1111-07, DIAL1128-07, DIAL1130-07, DIAL1138-07; BINs: BOLD:AAB5530, BOLD:ABZ0512, BOLD:ACE7697). There is a deep divergence between some of these sequences (maximum p-distance 3.21%) compared to the divergence between L. hudsoniellum and its nearest neighbour, L. nymphale (minimum p-distance 1.68%). No unique fixed substitutions distinguish L. hudsoniellum from all other western red-tailed L. ( Dialictus ).

Remarks

Lasioglossum hudsoniellum is undoubtedly the most common and widespread of the red-tailed L. ( Dialictus ). It occurs in all habitats from prairies to mountains to deserts. There is considerable variation in several characters over its range, which has led to much confusion with identification. Specimens from the easternmost part of its range in the Great Plains often have a black metasoma (even in the female), specimens from the western deserts often have pale wings with white hairs, and specimens from Texas and Mexico often have sparser punctures of the mesosoma and T1. In addition, there is significant variation in the length of the face, but this does not seem to be correlated with any other characters.

Given such extreme morphological, genetic, and geographic variation, L. hudsoniellum seems likely to include some cryptic species. However, clear consistencies between these sources of variation cannot be found to support a species hypothesis.

Figures

Fig. 1. Colour variation in the metasoma of Lasioglossum (D.) hudsoniellum (Cockerell, 1919). A. Yellow-orange. B. Red-orange. C. Reddish brown. D. Black. “Red-tailed” Lasioglossum (Dialictus) are considered to be any species with natural colour matching A or B.

Fig. 42. Georeferenced collection records of Lasioglossum (D.) hudsoniellum (Cockerell, 1919) (black squares) and predicted distribution by maximum entropy ecological niche modeling in Maxent (colour shading). Warmer colours indicate higher cloglog probability of occurrence.

Fig. 90. Male genitalia. A. Lasioglossum (D.) arenisaltans sp. nov. B. L. (D.) clastipedion sp. nov. C. L. (D.) clematisellum (Cockerell, 1904). D. L. (D.) hudsoniellum (Cockerell, 1919). E. L. (D.) julipile sp. nov. F. L. (D.) lilianae sp. nov. G. L. (D.) mesillense (Cockerell, 1898). H. L. (D.) meteorum sp. nov. I. L. (D.) miltolepoides sp. nov. J. L. (D.) minckleyi sp. nov. K. L. (D.) pallidellum (Ellis, 1914). L. L. (D.) pictum (Crawford, 1902). M. L. (D.) rufornatum sp. nov. N. L. (D.) spivakae sp. nov. O. L. (D.) testaceum (Robertson, 1897).P. L. (D.) vierecki (Crawford, 1904). Q. L. (D.) zephyrus (Smith, 1853). Genitalia are illustrated in dorsal view and positioned such that the gonostylus is parallel to the plane of vision. Retrorse lobes are also illustrated separately to the bottom left of each figure in lateral view, positioned parallel to the plane of vision.

Fig. 107. Pronotal angle (arrow and outline). A. Lasioglossum (D.) hudsoniellum (Cockerell, 1919), ♀, nearly 90 degrees. B. L. (D.) minckleyi sp. nov., ♀, obtuse. Scale bars: 0.5 mm.

Fig. 116. Metasomal sterna, ♂. A. Lasioglossum (D.) zephyrus (Smith, 1853), with hair concentrated in apicolateral areas of S3–5. B. Typical red-tailed Lasioglossum (Dialictus), with hair evenly distributed and S5 straight. C. L. (D.) hudsoniellum (Cockerell, 1919), with S5 concave.