Lasioglossum (Dialictus) clematisellum ( Cockerell, 1904 )

Gardner, Joel & Gibbs, Jason, 2020, The ‘ red-tailed’ Lasioglossum (Dialictus) (Hymenoptera: Halictidae) of the western Nearctic, European Journal of Taxonomy 725, pp. 1-242 : 58-69

Lasioglossum (Dialictus) clematisellum ( Cockerell, 1904)

Figs 28–30 , 90C , 91B , 106B , 114B , 117A

Halictus clematisellus Cockerell, 1904: 13 (holotype, ♀, deposited in USNM, examined).

Halictus clematisellus – Cockerell 1906: 429 (key).

Halictus (Chloralictus) clematisellus – Sandhouse 1924: 4 (key).

Lasioglossum (Chloralictus) clematisellum – Michener 1951: 1112 (catalog).

Dialictus clematisellus – Hurd 1979: 1965 (catalog). — Moure & Hurd 1987: 95 (catalog). Lasioglossum (Dialictus) clematisellum – Scott et al. 2011: 29 (checklist).

Diagnosis

Females of Lasioglossum clematisellum can be recognized by the characters for the L. clematisellum species complex (below), in addition to the metasomal terga lacking tomentum, the face and mesepisternum often duller than in other species in this group, scutum with most hairs relatively thin and weakly plumose and punctures of anteromedian area indistinct, and head and mesosoma blue-green with bright white hair. They are most similar to L. clastipedion sp. nov. and can only be separated with great difficulty, especially if specimens are worn. Females of L. clastipedion sp. nov. have the metasomal terga with at least some basolateral tomentum; this is the most reliable character. In addition, females of L. clastipedion sp. nov. often have the scutum with most hairs densely plumose and punctures of the anteromedian area distinct, and head and mesosoma more golden-green with slightly yellowish hair dorsally.

Males of L. clematisellum can be recognized by the characters for the Lasioglossum clematisellum species complex (below), in addition to the metasoma lacking tomentum, scutum with many hairs thin and weakly plumose, and frons densely punctate (i <1 pd). They are most similar to L. julipile sp. nov., L. perditum sp. nov., and L. clastipedion sp. nov. All of these species have the scutum with most hairs thick and densely plumose and metasomal terga with at least some basolateral tomentum. In addition, L. julipile sp. nov. and L. perditum sp. nov. have the frons more sparsely punctate (i=1–3 pd).

Etymology

Cockerell (1904) named this species after the plant genus Clematis , plus the Latin adjectival diminutive suffix - ellus.

Material examined

Holotype

UNITED STATES – New Mexico • ♀; Pecos ; [35.57° N, 105.68° W]; 14 Jul. 1903; W.P. Cockerell leg.; ex Clematis ligusticifolia ; USNM 12064 .

Other material

MEXICO – Nuevo Leon • 1 ♀; La Hediondilla, Galeana ; 25.0694° N, 100.7186° W; 1890 m a.s.l.; 29 Sep. 2013; LRF and CVM leg.; LRF. – Sonora • 1 ♀; 30 km E of Agua Prieta; 31.3097° N, 109.2542° W; 20 Apr. 2005; R.L. Minckley leg.; PCYU • 1 ♀; 30 km E of Agua Prieta; 31.3333° N, 109.26° W; 3 May 2001; A. Romero leg.; RLM • 1 ♀; 30 km E of Agua Prieta; 31.3333° N, 109.2597° W; 20 Aug. 2003; R.L. Minckley leg.; ex Baccharis sp.; PCYU • 1 ♀; 30 km E of Agua Prieta; 31.3133° N, 109.2503° W; 18 Jul. 2001; R. Minckley leg.; RLM • 1 ♀; 30 km E of Agua Prieta; 31.3222° N, 109.2772° W; 22 Jul. 2001; R. Minckley leg.; RLM • 1 ♀; 30 km E of Agua Prieta; 31.3186° N, 109.2508° W; 24 Jul. 2001; R. Minckley leg.; RLM • 1 ♀; 30 km E of Agua Prieta; 31.31° N, 109.2536° W; 4 Aug. 2001; R.L. Minckley leg.; RLM • 1♀; Municipio deAgua Prieta, Rancho Nogalitos ; 31.31177°N, 108.98622°W; 25 May 2008; RLM • 3 ♀♀; Rancho Puerta Blanca, Pollination Site 25 , Larrea, RPB, Loma Larrea ; 31.31389° N, 109.25028° W; 3 Apr. 2007; ex Salix gooddingii ; RLM • 1 ♀; Rancho San Bernardino ; 31.3133° N, 109.2503° W; 7 Aug. 2001; R.L. Minckley leg.; RLM • 2 ♀♀; Rancho San Bernardino ; 31.31° N, 109.2536° W; 25 Aug. 2001; R. Minckley leg.; RLM • 7 ♀♀; Rancho San Bernardino , 28 km E of Agua Prieta; [31.31° N, 109.25° W]; 6–7 Jul. 2000; R. Minckley et al. leg.; RLM • 2 ♀♀; same location as for preceding; 19 Jul. 2000; K. and R. Gyuro and S. Minckley leg.; RLM • 1 ♀; same location as for preceding; 22 Aug. 2000; R. Minckley and A. Romero leg.; RLM • 1 ♀; Rancho San Bernardino , 28 km E of Agua Prieta, Arroyo Grande ; [31.31° N, 109.25° W]; 14 Apr. 2000; R. Minckley leg.; ex Prosopis velutina ; RLM • 1 ♀; Rancho San Bernardino , Bosque dry , Site 4, Transect 1; 31.30992° N, 109.27783° W; 21 Jul. 2005; RLM • 6 ♀♀; Rancho San Bernardino , Bosque wet , Site 1; 31.33194° N, 109.27083° W; 10 Aug. 2006; RLM • 2 ♀♀; Rancho San Bernardino , Bosque wet , Site 7; 31.30972° N, 109.25417° W; 20 Apr. 2005; RLM • 1 ♀; Rancho San Bernardino , Bosque wet , Site 7, Transect 2; 31.31007° N, 109.25348° W; 22 Sep. 2001; RLM • 1 ♀; same location as for preceding; 12 Aug. 2002; RLM • 1 ♀; same location as for preceding; 1 Aug. 2004; RLM • 1 ♀; same location as for preceding; 25 Jul. 2007; RLM • 1 ♀; Rancho San Bernardino , Bosque wet , Site 8; 31.33333° N, 109.26528° W; 7 Apr. 2006; RLM • 1 ♀; Rancho San Bernardino , Cienega house ; 31.33098° N, 109.2676° W; 1 Apr. 2006; RLM • 1 ♀; Rancho San Bernardino , field, Site 1, Transect 1; 31.32422° N, 109.26663° W; 22 Jul. 2005; ex Convolvulus arvensis ; RLM • 1 ♀; same location as for preceding; 24 Jul. 2005; ex Convolvulus arvensis ; RLM • 1 ♀; Rancho San Bernardino , Los Ojitos , Lebuhn , Bee Diversity ; 31.30025° N, 109.2625° W; 19 Apr. 2002; RLM • 1 ♀; same location as for preceding; 3 May 2002; RLM • 1 ♀; Rancho San Bernardino , main ranch house ; 31.325° N, 109.265° W; 19 Jul. 2001; ex Lepidium thurberi ; RLM • 2 ♀♀; Rancho San Bernardino , Pollination Site 11, Sphaeralcea , 100 m SW of large dam on Silver Creek; 31.30972° N, 109.27778° W; 6 Aug. 2004; RLM • 1 ♀; same location as for preceding; 8 Aug. 2004; ex Chenopodium desiccatum ; RLM • 2 ♀♀; same location as for preceding; 8 Aug. 2004; ex Lepidium thurberi ; RLM • 1 ♀; same location as for preceding; 8 Aug. 2004; ex Salsola kali subsp. tragus ; RLM • 2 ♀♀; same location as for preceding; 10 Aug. 2004; ex Atriplex canescens ; RLM • 2 ♀♀; same location as for preceding; 10 Aug. 2004; ex Chenopodium neomexicanum ; RLM • 6 ♀♀; same location as for preceding; 10 Aug. 2004; ex Lepidium thurberi ; RLM • 1 ♀; Rancho San Bernardino , Pollination Site 12 , Physaria, Silver Creek , upstream of Big Dam; 31.3259° N, 109.28802° W; 13 Apr. 2006; ex Phacelia crenulata ; RLM • 1 ♀; same location as for preceding; 11 Apr. 2008; ex Physaria gordonii ; RLM • 1 ♀; same location as for preceding; 19 Apr. 2008; ex Descurainia sp.?; RLM • 1 ♀; Rancho San Bernardino , Pollination Site 21 , Sphaeralcea, Pond 100 m E of main gate; 31.31583° N, 109.26861° W; 25 Jul. 2007; ex Laennecia coulteri ; RLM • 2 ♀♀; Rancho San Bernardino , Pollination Site 24 , Physaria, Hay Hollow ; 31.31333° N, 109.25028° W; 5 Apr. 2007; ex Descurainia sp.?; RLM • 1 ♀; same location as for preceding; 5 Apr. 2007; ex Physaria gordonii ; RLM • 1 ♀; same location as for preceding; 5 Apr. 2007; ex Salix gooddingii ; RLM • 2 ♀♀; same location as for preceding; 14 Apr. 2007; ex Physaria gordonii ; RLM • 1 ♀; Rancho San Bernardino , Pollination Site 27 , Horse Corral , 70 m N of main ranch house; 31.32556° N, 109.26667° W; 8 Aug. 2007; ex Lepidium thurberi ; RLM • 1 ♀; Rancho San Bernardino , Pollination Site 6 , Sphaeralcea , field 1; 31.325° N, 109.26361° W; 1 Aug. 2005; ex Convolvulus arvensis ; RLM • 1 ♀; Rancho San Bernardino , quarry, at road crossing; 31.32042° N, 109.2681° W; 2 Sep. 2003; RLM • 1 ♀; Rancho San Bernardino , quarry, general area; 31.3195° N, 109.27028° W; 8 May 2001; ex Phacelia sp.?; RLM • 2 ♀♀; Rancho San Bernardino , quarry, gravel pit; 31.31964° N, 109.26926° W; 18 Aug. 2003; RLM • 1 ♀; Rancho San Bernardino , roadside E of quarry; 31.31996° N, 109.26848° W; 20 Aug. 2003; RLM • 3 ♀♀; Rancho San Bernardino , roadside W of quarry; 31.31948° N, 109.2701° W; 18 Aug. 2003; RLM • 2 ♀♀; same location as for preceding; 19 Aug. 2003; RLM • 1 ♀; Rancho San Bernardino ; 31.32907° N, 109.26553° W; 1 Aug. 2000; RLM • 1 ♀; same location as for preceding; 30 Jul. 2005; ex Lepidium thurberi ; RLM • 1 ♀; Rancho San Bernardino , Riparian 3; 31.31389° N, 109.25028° W; 20 Aug. 2003; RLM • 1 ♀; same location as for preceding; 25 Aug. 2003; RLM • 1 ♀; Rancho San Bernardino , Riparian , Site 1; 31.33333° N, 109.25972° W; 20 Aug. 2003; ex Baccharis sarothroides ; RLM • 1 ♀; same location as for preceding; 20 Aug. 2003; RLM • 1 ♀; same location as for preceding; 7 Apr. 2006; RLM • 2 ♀♀; same location as for preceding; 13 May 2006; RLM • 1 ♀; Rancho San Bernardino , Riparian , Site 2; 31.33194° N, 109.25833° W; 7 Apr. 2006; RLM • 1 ♀; Rancho San Bernardino , Riparian , Site 3, Transect 1; 31.31362° N, 109.25468° W; 18 Sep. 2001; RLM • 1 ♀; Rancho San Bernardino , Riparian , Site 3, Transect 3; [31.32° N, 109.26° W]; 25 Sep. 2001; RLM • 1 ♀; Rancho San Bernardino , Riparian , Site 4, Transect 1; 31.32965° N, 109.25587° W; 18 Sep. 2001; RLM • 2 ♀♀; Rancho San Bernardino , Silver Creek , Big Dam , Pol 12; 31.3259° N, 109.28802° W; 9 May 2006; RLM • 1 ♀; Rancho San Bernardino , Tamarisk Site 3, Rio San Bernardino ; 31.30743° N, 109.25687° W; 16 Jun. 2004; RLM • 2 ♀♀; Rancho San Bernardino , Tamarisk Site 5 , Rio San Bernardino ; 31.3071° N, 109.25682° W; 12 Jun. 2004; ex Tamarix ramosissima ; RLM • 1 ♀; Rancho San Bernardino , Tamarisk , at Rio San Bernardino; 31.30725° N, 109.25675° W; 26 May 2004; ex Tamarix ramosissima ; RLM • 1 ♀; same location as for preceding; 29 May 2004; ex Tamarix ramosissima ; RLM • 1 ♀; same location as for preceding; 3 Jun. 2004; ex Tamarix ramosissima ; RLM • 1 ♀; Rancho San Bernardino , Tamarisk , at Silver Creek; 31.31611° N, 109.25806° W; 26 May 2004; ex Tamarix ramosissima ; RLM. – Zacatecas • 1 ♀, 3 ♂♂; 45 mi. SW of Saltillo; [24.7° N, 101.3° W]; 14 Sep. 1977; R.W. Brooks leg.; SEMC.

UNITED STATES – Arizona • 1 ♀; Pinal Co., ~ 24 km S of Mammoth; 32.5558° N, 110.5164° W; 30 Apr. 2011; R. Laport leg.; ex Larrea tridentata ; RLM • 1 ♀; Cochise Co., 18 km W of Portal; [31.91° N, 109.33° W]; 18 Jun. 1993; R.L. Minckley leg.; SEMC • 1 ♀; Cochise Co., 2 mi. NE of Portal; [31.94° N, 109.11° W]; 30 Jul. 1959; Statham leg.; AMNH • 1 ♀; 21 mi. SW of Prescott; [34.32° N, 112.73° W]; 17 Jul. 1950; C.D. Michener leg.; ex Prosopis juliflora ; SEMC • 1 ♀; 23 mi. W of Winslow; [35.11° N, 111.09° W]; 3 Jul. 1953; E.S. Ross leg.; CAS • 1 ♀; Cochise Co., 3.2 km E of Apache; [31.69° N, 109.1° W]; 4–11 Jun. 1987; B.N. Danforth leg.; ex Senecio flaccidus ; SEMC • 1 ♀; Cochise Co., 4 mi. E of Willcox; [32.25° N, 109.76° W]; 18 Aug. 1986; R.J. McGinley and B.M. Norden leg.; USNM • 1 ♀; Graham Co., 4 mi. W of Calva; [33.19° N, 110.25° W]; 17 Jun. 1956; J.W. MacSwain and D.D. Linsdale leg.; ex Prosopis sp.; EMEC • 1 ♀; Cochise Co., 6 mi. S of Willcox; [32.16° N, 109.83° W]; 23 Aug. 1986; R.J. McGinley and B.M. Norden leg.; USNM • 1 ♀; Pima Co., 7 km E of Silverbell; 32.3667° N, 111.4333° W; 732 m a.s.l.; 4 Apr. 1994; R.L. Minckley and S.G. Reyes leg.; ex Sphaeralcea sp.; SEMC • 1 ♀; Pima Co., 7 mi. N of Tucson; [32.33° N, 110.97° W]; 4 Sep. 1968; D.R. Miller and J.E. Lauck leg.; UCDC • 1 ♀; Cochise Co., 8 mi. NE of Portal; [32° N, 109.48° W]; 9 May 1956; M. Statham leg.; AMNH • 2 ♂♂; Cochise Co., 8.1 km NE of Douglas; [31.4° N, 109.49° W]; 13 Jun. 1987; B.N. Danforth leg.; ex Lepidium ; SEMC • 1 ♂; Cochise Co., Apache ; [31.69° N, 109.13° W]; 8 Aug. 1972; G. and K. Eickwort leg.; CUIC • 5 ♀♀, 1 ♂; Cochise Co., Apache vicinity ; [31.69° N, 109.13° W]; 7 Jun. 1987; B.N. Danforth leg.; ex Lepidium ; SEMC • 7 ♀♀, 15 ♂♂; Arivaca ; [31.58° N, 111.33° W]; 13 Jul. 1950; R.H. Beamer leg.; SEMC • 1 ♂; same location as for preceding; 13 Jul. 1950; J.G. Rozen leg.; SEMC • 1 ♂; Arivaipa ; [32.96° N, 110.36° W]; 25 Aug. 1933; Bryant leg.; CAS • 1 ♀; Cochise Co., Benson ; [31.97° N, 110.29° W]; 14–15 Sep. 1974; E.G. and J.M. Linsley leg.; ex Helianthus annuus ; EMEC • 4 ♀♀; Carrizo Creek ; [33.9° N, 110.28° W]; 16 Jun. 1950; R.H. Beamer leg.; ex Euphorbia albomarginata ; SEMC • 1 ♀; same location as for preceding; 16 Jun. 1950; L.D. Beamer leg.; ex Sapindus drummondii ; SEMC • 1 ♀; Granite Dells ; [34.61° N, 112.41° W]; 3 Jul. 1950; H. Wright leg.; SEMC • 1 ♀; Pima Co., Madera Canyon , 13 km S and 10 km E of Continental; 31.7488° N, 110.8872° W; 20 Jul. 2010; R.E. Wrigley leg.; WRME • 1 ♀; Mule Pass no. Bisbee ; [31.458° N, 109.942° W]; 24 Mar. 1946; C.D. Michener leg.; ex yellow composite; SEMC • 2 ♀♀; Cochise Co., nr Willcox Blue Sky Rd.; 32.23199° N, 109.77807° W; 1279 m a.s.l.; 14 Aug. 2007; Gibbs and Sheffield leg.; PCYU • 2 ♀♀; Patagonia ; [31.54° N, 110.76° W]; 10 Sep. 1957; T.R. Haig leg.; UCDC • 1 ♀; Cochise Co., Portal vicinity ; [31.91° N, 109.14° W]; 2–12 Jun. 1987; B.N. Danforth leg.; ex Prosopis ; SEMC • 1 ♀; Pinal Co., 12 mi. N of Redington; [32.6° N, 110.5° W]; 22 May 1975; J.L. Neff leg.; ex Prosopis velutina ; CTMI • 1 ♀; same location as for preceding; 22 May 1975; J.L. Neff leg.; ex Larrea tridentata ; CTMI • 1 ♀, 1 ♂; Cochise Co., S of Benson; 31.935578° N, 110.28643° W; 1273 m a.s.l.; 14 Aug. 2007; Gibbs and Sheffield leg.; PCYU • 1 ♀; Salt River Canyon ; [33.7° N, 110.8° W]; 16 Jun. 1950; R.H. Beamer leg.; SEMC • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3386° N, 109.2719° W; 19 Apr. 2005; R.L. Minckley leg.; PCYU • 1 ♀; Cochise Co., San Bernardino NWR ; 31.3417° N, 109.2583° W; 30 Mar.2002; A.Romero leg.; RLM • 1♀; same location as for preceding; 28Apr.2004; R.L.Minckley leg.; RLM • 1 ♀; Cochise Co., San Bernardino NWR ; 31.3344° N, 109.2681° W; 1 Sep. 2001; R.L. Minckley leg.; RLM • 1 ♀; Cochise Co., San Bernardino NWR ; 31.3406° N, 109.2575° W; 8 Sep. 2001; R.L. Minckley leg.; RLM • 1 ♀; Cochise Co., San Bernardino NWR ; 31.3417° N, 109.2583° W; 26 Jul. 2001; R.L. Minckley leg.; RLM • 2 ♀♀; Cochise Co., San Bernardino NWR , Bosque dry , Site 6; 31.34058° N, 109.25753° W; 6 Aug. 2003; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Bosque dry , Site 6, Transect 2; 31.34058° N, 109.25747° W; 19 Jul. 2005; RLM • 1 ♀; same location as for preceding; 16 Aug. 2005; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Bosque dry , Site 7; 31.34118° N, 109.25813° W; 6 Aug. 2003; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Bosque dry , Site 7, Transect 2; 31.34168° N, 109.25825° W; 11 Oct. 2002; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Bosque wet , Site 2, Transect 2; 31.33627° N, 109.26752° W; 15 Aug. 2005; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Bosque wet , Site 4, Transect 1; 31.33693° N, 109.24361° W; 11 Aug. 2005; RLM • 1 ♀; Cochise Co., San Bernardino NWR , field, Site 5 , Transect 3; 31.33417° N, 109.26778° W; 30 Jul. 2004; RLM • 4 ♀♀; Cochise Co., San Bernardino NWR , field, Site 6; 31.33867° N, 109.2719° W; 8 Aug. 2003; RLM • 1 ♀; Cochise Co., San Bernardino NWR , field, Site 6, Transect 4; 31.33444° N, 109.26806° W; 30 Jul. 2004; RLM • 1 ♀; Cochise Co., San Bernardino NWR , field, Site 8; 31.33765° N, 109.26068° W; 9 May 2008; RLM • 2 ♀♀; Cochise Co., San Bernardino NWR , field, Site 8, Transect 2; 31.3381° N, 109.2605° W; 30 Jul. 2004; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Grassland , Site 2, Transect 3; 31.3358° N, 109.2702° W; 30 Jul. 2004; RLM • 2 ♀♀; Cochise Co., San Bernardino NWR , Grassland , Site 2, Transect 4; 31.3358° N, 109.2702° W; 11 Aug. 2005; RLM • 6 ♀♀; Cochise Co., San Bernardino NWR , Grassland , Site 4; 31.33495° N, 109.25314° W; 5 Aug. 2003; RLM • 1 ♀; same location as for preceding; 11 Aug. 2005; RLM • 2 ♀♀; Cochise Co., San Bernardino NWR , Grassland , Site 4, Transect 2; 31.335° N, 109.25389° W; 31 Jul. 2004; RLM • 2 ♀♀; Cochise Co., San Bernardino NWR , Grassland , Site 5 ; 31.33683° N, 109.25308° W; 5 Aug. 2003; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Riparian , Site 5 , Transect 1; 31.33448° N, 109.25245° W; 15 Oct. 2002; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Robertson cienega ; 31.33417° N, 109.25222° W; 19 Aug. 2001; ex Xanthisma sp.?; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Shrub , Site 5 , Transect 2; 31.34258° N, 109.27013° W; 27 Jul. 2007; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Shrub , Site 8; 31.34167° N, 109.25278° W; 5 Aug. 2003; RLM • 1 ♀; Cochise Co., San Bernardino NWR , Shrub , Site 8, Transect 3; 31.34072° N, 109.253° W; 11 Aug. 2005; RLM • 1 ♀; Cochise Co., San Simon ; [32.27° N, 109.23° W]; 23 Apr. 1961; Rozen and Schrammel leg.; AMNH • 7 ♂♂; Pima Co., Silver Bell Bajada , IBP Desert Scrub Site ; [32.4° N, 111.5° W]; J.L. Neff leg.; CAS • 1 ♂; same location as for preceding; 1 Sep. 1974; J.L. Neff leg.; ex Trianthema portulacastrum ; CUIC • 1 ♀; Cochise Co., St David Cienega ; 31.8465° N, 110.2275° W; 4 Apr. 2016; C.M. Delphia and J.B. Runyon leg.; MTEC • 1 ♂; Cochise Co., Tombstone ; [31.71° N, 110.07° W]; 3–5 Jul. 1971; M.L. Lindsey leg.; CUIC • 2 ♀♀; Tucson ; [32.22° N, 110.97° W]; 4 Mar. 1934; Bryant leg.; CAS • 3 ♀♀; same location as for preceding; 6 Jul. 1950; R.H. Beamer leg.; ex Prosopis glandulosa ; SEMC • 21 ♀♀; White Mts; [33.9° N, 109.6° W]; 19 Jun. 1950; R.H. Beamer leg.; ex Amorpha fruticosa ; SEMC • 1 ♂; same location as for preceding; 19 Jun. 1950; L.D. Beamer leg.; ex Penstemon superbus ; SEMC • 1 ♀; Willcox ; [32.25° N, 109.83° W]; 2005; L. Packer leg.; PCYU • 1 ♀; Cochise Co., 2 mi. SE of Willcox; 32.227° N, 109.781° W; 24 Aug. 2016; J.L. Neff leg.; ex Eriogonum abertianum ; CTMI • 1 ♀; Cochise Co., 6.4 km SE of Willcox on Blue Sky Rd; 32.2275° N, 109.7814° W; 1250 m a.s.l.; 21 Aug. 2001; C. Michener leg.; ex Psorothamnus scoparius ; SEMC. – California • 1 ♀; Riverside Co., 3 mi. E of Moreno Valley, Redlands Boulevard ; [33.94° N, 117.16° W]; 2 Jul. 1982; N.J. Smith leg.; UCDC • 1 ♀; Los Angeles Co., Frenchmans Flats ; [34.617° N, 118.745° W]; 8 Jun. 1966; R.J. Hamton leg.; LACM • 6 ♀♀; Los Angeles Co., 2 mi. W of Pearblossom; [34.51° N, 117.94° W]; 12 May 1956; E.G. Linsley leg.; EMEC • 1 ♀; Los Angeles Co., Pico Canyon , 2 mi. W of Newhall; [34.38° N, 118.6° W]; 12 Aug. 1975; J. Lyon leg.; ex Eriogonum fasciculatum ; LACM • 3 ♀♀, 22 ♂♂; Riverside ; [33.97° N, 117.32° W]; 1953; Timberlake leg.; ex Alyssum ; EMEC. – Colorado • 1 ♂; Montrose Co., 18 mi. SE of Gateway; [38.49° N, 108.75° W]; 19 Aug. 1963; N. and B. Marston leg.; AMNH • 3 ♀♀, 5 ♂♂; Fremont Co., 6 mi. NE of Florence; [38.45° N, 105.04° W]; 11Aug. 1964; J.G., B.L. and K.C. Rozen leg.; AMNH • 1 ♀; Boulder; [40.02° N, 105.27° W]; 10 Aug. 1935; C.D. Michener leg.; ex Cleome ; SEMC • 1 ♀; Bent Co., John Martin Reservoir ; [38.1° N, 103° W]; 6 Jul. 1978; G. E. Bohart leg.; ex Melilotus ; WRME • 1 ♂; Las Animas Co., Model ; 37.493° N, 103.8345° W; 23 Jul. 2009; J. Newton leg.; PCYU • 1 ♀; Las Animas Co., Pergatoire Riv. Canyon , dinosaur track site , Comanche Nat. Grasslands ; [37.617° N, 103.598° W]; 21 Jun. 2005; H. Guarisco leg.; ex Dalea candida var. oligophylla ; SEMC • 1 ♀; Delta Co., R. M. R. C. ; [39° N, 108° W]; 3 Jul. 1996; R. Konold leg.; ex buckwheat; CSUC • 1 ♀; same location as for preceding; 3 Jul. 1996; R. Konold leg.; ex Alyssum ; CSUC • 1 ♀; same location as for preceding; 23 Jun. 1996; R. Konold leg.; ex Alyssum ; CSUC • 1 ♀; White Rocks , Boulder ; [40.06° N, 105.16° W]; 8 Jul. 1935; C.D. Michener leg.; ex Dalea ; SEMC. – Montana • 1 ♀; Gallatin Co., Gallatin Grown ; 45.7713° N, 111.2953° W; 25 Jul. 2013; C.M. Delphia leg.; MTEC • 1 ♀; same location as for preceding; 30 Aug. 2013; C.M. Delphia leg.; MTEC • 2 ♀♀; same location as for preceding; 24 Jul. 2015; C.M. Delpha and J.L. Monte leg.; MTEC • 2 ♀♀; same location as for preceding; 9 Jul. 2015; C.M. Delpha and J.L. Monte leg.; MTEC • 3 ♀♀; same location as for preceding; 2 Aug. 2013; C.M. Delphia leg.; MTEC • 1 ♀; same location as for preceding; 8Aug. 2013; C.M. Delphia leg.; MTEC • 1 ♀; Gallatin Co., Towne’s Harvest Garden , 2730 W of Garfield Avenue; [45.66° N, 111.07° W]; 8 Aug. 2011; C.M. Delphia leg.; MTEC. – Nevada • 1 ♀; Lincoln Co., Caliente ; [37.62° N, 114.51° W]; 14 Jul. 19?8; F.D. Parker leg.; UCDC • 1 ♀; Lincoln Co., Kershaw Ryan State Park ; [37.59° N, 114.52° W]; 14 Jul. 1971; J. Doyen leg.; EMEC. – New Mexico • 1 ♂; Hidalgo Co., 10 mi. S of Animas; [31.8° N, 108.81° W]; 22 Aug. 1972; G. C. Eickwort leg.; CUIC • 1 ♀; Hidalgo Co., 13 mi. N of Rodeo; [32.02° N, 109.03° W]; 1372 m a.s.l.; 16 Aug. 1964; C.D. Michener leg.; ex Pectis ; SEMC • 1 ♀; same location as for preceding; 18 Aug. 1964; C.D. Michener leg.; ex Wislizenia refracta ; SEMC • 1 ♀, 5 ♂♂; Hidalgo Co., 14 mi. S of Animas; [31.74° N, 108.81° W]; 13 Jul. 1987; B. Danoff leg.; ex Lepidium ; SEMC • 1 ♀; Hidalgo Co., 15 km N of Rodeo; [31.97° N, 109.03° W]; 1300 m a.s.l.; 6 Jul. 1993; Robert L. Minckley leg.; ex Glandularia bipinnatifida ; SEMC • 1 ♀; Hidalgo Co., 15 mi. S of Animas; [31.73° N, 108.81° W]; 24 Aug. 1986; R.J. McGinley and B.M. Norden leg.; USNM • 1 ♀; Hidalgo Co., 3.5 mi. SW of Rodeo; [31.8° N, 109.07° W]; 16 Aug. 1986; R.J. McGinley and B.M. Norden leg.; USNM • 1 ♀; Alamogordo ; [32.9° N, 105.96° W]; 13 May 1902; ANSP • 1 ♀; same location as for preceding; 26 Jun. 1940; L.J. Lipovsky leg.; SEMC • 1 ♀; Hidalgo Co., Animas; [31.95° N, 108.81° W]; 9 May 1993; R.L. Minckley and K. Toal leg.; SEMC • 1 ♀; Hidalgo Co., 16 mi. S of Animas; 31.7213° N, 108.8224° W; 29 Aug. 2013; T.L. Griswold leg.; ex Baccharis salicina ; BBSL • 1 ♀; Hidalgo Co., Animas , 8.8 km E of junc. Sock N Shoe Rd; 31.9572° N, 108.7144° W; 1390 m a.s.l.; 15 Aug. 2000; R. Brooks leg.; ex Fallugia paradoxa ; SEMC • 1 ♀; Hidalgo Co., Antelope Corral , 6 mi. N of Rodeo; 31.9167–31.9333° N, 109–109.0166° W; 1231 m a.s.l.; 25 Jun. 1984; H.A. Hespenheide leg.; ex Artemisia / Atriplex ; LACM • 1 ♀; Hidalgo Co., 12.8 mi. W of Hachita; 31.965° N, 108.6111° W; 1440 m a.s.l.; 27 Sep. 1999; R. Brooks and C. Michener leg.; ex Heliomeris multiflora var. nevadensis ; SEMC • 1 ♂; Socorro Co., Bosque del Apache ; [33.8° N, 106.89° W]; 1375 m a.s.l.; 15 Jul. 1991; Ilan Yarom leg.; SEMC • 19 ♀♀; Chaves Co., Bottomless Lakes , 10 mi. E of Roswell; [33.34° N, 104.34° W]; 20 May 1969; Brothers, Krueger and Michener leg.; SEMC • 1 ♂; Eddy Co., Carlsbad ; [32.42° N, 104.23° W]; 20 May 1969; Brothers, Krueger and Michener leg.; ex Lepidium integrifolium ; SEMC • 2 ♀♀; Carrizozo; [33.64° N, 105.88° W]; 10 Jun. 1950; L.D. Beamer leg.; ex Helenium laciniatum ; SEMC • 9 ♂♂; Luna Co., Columbus ; [31.83° N, 107.64° W]; 21 Jul. 1982; W.J. Pulawski leg.; CAS • 1 ♂; same location as for preceding; 21 Jul. 1982; W.J. Pulawski leg.; CUIC • 1 ♀; Highrolls ; [32.95° N, 105.84° W]; 29 May 1902; ANSP • 1 ♀; Hot Springs ; [35.65° N, 105.29° W]; 22 Jul. 1959; W.J. Arnold leg.; SEMC • 1 ♀; Jemez Springs .; [35.77° N, 106.69° W]; 1 Jul. 1941; R.H. Beamer leg.; SEMC • 2 ♀♀; Nogal ; [33.56° N, 105.71° W]; 21 Jul. 1952; C. Liang, W. LaBerge, R.H. and L.D. Beamer leg.; SEMC • 5 ♀♀; Eddy Co.; 32.1095° N, 104.4716° W; 1115 m a.s.l.; 19 Jul. 2010; J.D. Herndon leg.; ex Plantago sp.; BBSL • 1 ♀; Hidalgo Co., S. of Animas; [31.9° N, 108.81° W]; 2003; L. Packer leg.; PCYU. – Texas • 5 ♀♀; Jeff Davis Co., 0.5 mi. S of Kent; [31.06° N, 104.22° W]; 29 May 1999; L. Packer leg.; PCYU • 1 ♀; Reeves Co., 3 mi. NW of Pecos; 31.453° N, 103.508° W; 6 Jul. 2007; J.L. Neff and A. Hook leg.; ex Lepidium alyssoides ; CTMI • 1 ♀; Santa Elena Canyon , Big Bend National Park ; [29.165° N, 103.612° W]; 26 Aug. 1967; R.C. Gardner and C.R. Kovacic leg.; UCDC. – Utah • 1 ♀; Kane Co., 1.0 mi. E of Nipple Spring; 37.189° N, 111.5368° W; 16 May 2002; K. Fordney leg.; ex Tamarix sp.; BBSL • 3 ♀♀; Garfield Co., 10 mi. N of Escalante; [37.9° N, 111.6° W]; 19 Jun. 2000; F.D. Parker leg.; BBSL • 10 ♀♀; Blanding ; [37.62° N, 109.48° W]; 23 May 1967; G. F. Knowlton leg.; BBSL • 18 ♀♀; San Juan Co., Blanding ; [37.62° N, 109.48° W]; 23 May 1967; G.F. Knowlton leg.; UCDC • 1 ♀; Kane Co., Butler Valley Dam ; 37.4693° N, 111.871° W; 19 Jul. 2002; J. Richmond leg.; ex Helianthus petiolaris ; PCYU • 2 ♀♀; Castle Valley ; [38.64° N, 109.41° W]; 10 May 1949; G.F. Knowlton leg.; SEMC • 1 ♀; Colorado River , 4 mi. N of Moab; [38.62° N, 109.51° W]; 1 Aug. 1987; R.S. Beal leg.; CSUC • 1 ♀; Colorado River , 5 mi. N of Moab; [38.63° N, 109.49° W]; 1 Aug. 1987; R.S. Beal leg.; CSUC • 2 ♀♀; Emery Co., Green River ; 38.9885° N, 110.162° W; 9 May 2000; F.D. Parker leg.; BBSL • 1 ♀; Kane Spgs ., 15 mi. SE of Moab; [38.392° N, 109.452° W]; 9 Aug. 1957; C.D. Michener leg.; ex Cleome lutea ; SEMC • 1 ♀, 9 ♂♂; Grand Co., Moab Hwy 128, 2 mi marker roadside; [38.6117° N, 109.5513° W]; 15 Aug. 1992; L. Packer leg.; PCYU • 2 ♀♀; Uintah Co., Ouray Valley ; [40° N, 109.5° W]; 12 Jul. 1956; G.F. Knowlton leg.; SEMC • 2 ♀♀; Garfield Co., Pine Creek , 4.5 mi. N of Escalante; 37.9031° N, 111.6686° W; 19 Jun. 2000; J. Grixti leg.; ex Tamarix sp.; BBSL • 3 ♀♀; same location as for preceding; 19 Jun. 2000; R. Sellars leg.; ex Tamarix sp.; BBSL • 2 ♀♀; same location as for preceding; 19 Jun. 2000; O. Messinger leg.; ex Tamarix sp.; BBSL • 4 ♀♀; Uintah Co., Vernal ; [40.46° N, 109.53° W]; 18 Jul. 1961; G.F. Knowlton leg.; UCDC. – Wyoming • 1 ♀; Uinta Co., Lonetree ; [41.05° N, 110.15° W]; 17 Jul. 1961; G.F. Knowlton leg.; UCDC .

Redescription

Female

COLOURATION. Head and mesosoma blue-green to golden-green; clypeus apical colour reddish brown to orange; labrum reddish brown to orange; mandible orange with black basal spot or band and red tip; flagellum black to reddish brown dorsally, reddish brown to orange ventrally; pronotal lobe reddish brown to orange; metasoma red to orange with dark spiracular spots on T3–4; legs reddish brown with femur-tibia joints, tibia apices, and tarsi mostly orange; tegula pale amber; wing membrane hyaline, veins with subcosta dark brown, otherwise pale amber.

PUBESCENCE. Body hair colour white. Tomentum dense on paraocular area, gena, pronotal collar and lobe, space between pronotal lobe and tegula, and metanotum anterior margin; sparse on clypeus basal half, supraclypeal area, preëpisternum, metepisternum, and T4 basally. Scutum hair thin to slightly densely plumose. Wing hairs light, very short and sparse. Acarinarial fan complete or nearly so, sparse. T2 fringes very sparse, T3 fringes very sparse.

SURFACE SCULPTURE. Clypeus punctures dense in basal half (i <1 pd), large and irregularly spaced apically (i <2 pd), sculpture shiny, sometimes weakly tessellate basally; supraclypeal area punctures sparse (i=1–4 pd), becoming dense laterally (i ≤ 1 pd), sculpture shiny, sometimes becoming weakly tessellate laterally; paraocular area punctures dense (i ≤ 1 pd), sometimes crowded around antenna socket, sculpture shiny, sometimes weakly tessellate around antenna socket; frons punctures dense (i <1 pd), sculpture tessellate; vertex punctures fine, dense laterally (i ≤ 2 pd), moderately sparse medially (i =1–2 pd), sculpture shiny or weakly tessellate; gena punctures moderately sparse (i=1–3 pd), sculpture shiny; postgena sculpture tessellate and sometimes weakly rugulose; tegula punctures absent; scutum punctures moderately dense (i ≤ 2 pd), becoming sparser submedially (i=1–3 pd) and denser posteromedially (i <1 pd), sculpture shiny to weakly tessellate, becoming more strongly tessellate anteromedially; scutellum punctures moderately dense (i ≤ 2 pd), becoming sparser submedially (i =1–3 pd) and denser posteromedially (i <1 pd), diversopunctate, sculpture shiny, sometimes becoming tessellate marginally and on median line; metanotum sculpture shiny and finely, densely punctate (i <1 pd), becoming rugulose laterally; metapostnotum rugae strong, highly anastomosing, reaching margin, sculpture tessellate; preëpisternum sculpture areolate; hypoepimeron punctures dense (i <1 pd), sculpture shiny or weakly imbricate; mesepisternum punctures moderately dense (i =1–2 pd), becoming dense ventrally (i <1 pd), sculpture shiny or weakly imbricate; metepisternum sculpture rugulosolineate dorsally, areolate or tessellate and densely punctate (i <1 pd) ventrally; propodeum lateral face sculpture tessellate; propodeum posterior face sculpture tessellate; T1 anterior face sculpture shiny; T1 dorsal surface punctures moderately sparse (i= 1–4 pd), becoming finer or obscure on rim and absent in large apicolateral oval patches, sculpture shiny; T2 disc punctures moderately dense (i=1–3 pd), disc sculpture shiny, rim punctures very fine, sometimes obscure, sparse (i = 2–4 pd), rim sculpture weakly coriarious.

STRUCTURE. Face length/width ratio 0.83 (± 0.02 SD). Clypeus projecting ~67% below suborbital tangent; clypeal area length/width ratio 0.41 (± 0.04 SD); apicolateral denticles low rounded knobs; supraclypeal area length/width ratio 0.67 (± 0.08 SD). Forewing with 3 submarginal cells; pronotal angle obtuse; tegula shape normal. Intertegular distance 0.86 (± 0.03 SD) mm. Scutum length/width ratio 0.78 (± 0.02 SD); scutum/scutellum length ratio 2.99 (± 0.16 SD); scutellum/metanotum length ratio 1.51 (± 0.13 SD); metanotum/metapostnotum length ratio 0.65 (± 0.06 SD). Propodeum lateral carinae not reaching dorsal margin; oblique carina very weak or absent. T2 depressed apical rim length less than 50% of segment. (n =11)

Male

COLOURATION. Head and mesosoma blue to olive green; clypeus apical colour black to reddish brown; labrum reddish brown to orange; mandible orange with black basal spot and red tip; flagellum reddish brown dorsally, orange ventrally; pronotal lobe reddish brown; metasoma black to dark reddish brown with rims of terga and sterna broadly translucent reddish brown and downcurved lateral areas of terga becoming translucent orange; legs reddish brown with femur-tibia joints, base and apex of tibiae, and tarsi orange; tegula orange; wing membrane hyaline, veins dark to light brown.

PUBESCENCE. Body hair colour white. Tomentum dense on face below eye emargination, gena, pronotal angle and lobe, and space between pronotal lobe and tegula; sparse on episterna. Scutum hair thin to moderately plumose. Sterna hair short (0.75–1.25 OD), densely plumose, dense and erect. Wing hairs dark or light, short and sparse.

SURFACE SCULPTURE. Clypeus punctures dense to moderately dense (i ≤ 2 pd), sculpture shiny; supraclypeal area punctures dense to moderately dense (i ≤ 2 pd), sculpture shiny; paraocular area punctures dense (i <1 pd), sculpture shiny; frons punctures crowded (i=0 pd), sculpture shiny to weakly imbricate; vertex punctures moderately dense laterally (i=1–2 pd), sparse medially (i =1–3 pd), sculpture shiny to imbricate; gena punctures fine, sparse (i= 1–3 pd), obscure, sculpture shiny, becoming weakly ruguloso-lineate posteriorly and ventrally; postgena sculpture imbricate to ruguloso-lineate; tegula punctures absent; scutum punctures dense to moderately sparse (i ≤ 2 pd), becoming sparse submedially (i =1–3 pd), sculpture shiny to weakly tessellate, becoming more strongly tessellate anteromedially; scutellum punctures dense marginally and on median line (i ≤ 1 pd), sparse submedially (i= 1–3 pd), sculpture shiny; metanotum sculpture shiny and densely to moderately sparsely punctate (i ≤ 2 pd); metapostnotum rugae strong, anastomosing, reaching margin, sculpture shiny to weakly imbricate; preëpisternum sculpture shiny with crowded punctures (i=0 pd); hypoepimeron punctures dense (i ≤ 1 pd), sculpture shiny; mesepisternum punctures dense (i ≤ 1 pd), sculpture shiny; metepisternum sculpture lineate dorsally, areolate-rugose ventrally; propodeum lateral face punctures obscure, sculpture imbricate to rugulose; propodeum posterior face sculpture rugulose; T1 anterior face sculpture shiny; T1 dorsal surface punctures dense (i ≤ 1 pd), becoming sparse towards rim (i=1–3 pd) and absent or very sparse in small apicolateral oval patches, sculpture shiny; T2 disc punctures dense (i ≤ 1 pd), becoming sparse toward premarginal line (i=1–3 pd), disc sculpture shiny, rim punctures sparse medially (i =1–4 pd), absent laterally, rim sculpture shiny.

STRUCTURE. Face length/width ratio 0.83 (± 0.02 SD). F1:pedicel length ratio 0.89 (± 0.13 SD); F2:F1 length ratio 1.97 (± 0.23 SD); F2 length/width ratio 1.56 (± 0.12 SD); F9 length/width ratio 1.39 (± 0.09 SD). Forewing with 2 or 3 submarginal cells; pronotal angle obtuse; tegula shape normal. Intertegular distance 0.75 (± 0.05 SD) mm. Scutum length/width ratio 0.89 (± 0.05 SD); scutum/ scutellum length ratio 2.77 (± 0.14 SD); scutellum/metanotum length ratio 1.68 (± 0.17 SD); metanotum/ metapostnotum length ratio 0.66 (± 0.03 SD). Propodeum lateral carinae not reaching dorsal margin; oblique carina absent. (n=10)

GENITALIA. As in Fig. 90C . Gonocoxite relatively narrow, rounded. Gonostylus with a few long hairs. Retrorse lobe relatively narrow, ovoid, covered in sparse short hairs.

Range

Southwestern Montana south to Zacatecas and west to California ( Fig. 30 ).

Floral records

AIZOACEAE Martinov : Trianthema L. T. portulacastrum L. • AMARANTHACEAE Juss. Atriplex L. A. canescens (Pursh) Nutt. • Chenopodium L. C. desiccatum A. Nelson • C. neomexicanum Standl. • Salsola L. S. kali L. S. k. subsp. tragus (L.) Čelak. • ANACARDIACEAE R.Br. Schinus L. S. molle L. (MH87) • APIACEAE Lindl. Apium L. A. graveolens L. (MH87) • ASPARAGACEAE Juss. Nolina Michx. (MH87) • ASTERACEAE Giseke : Baccharis : B. salicina Torr. & A.Gray • B. sarothroides A. Gray • Chrysothamnus (MH87) • Ericameria : E. palmeri (A.Gray) H. M. Hall (MH87) • Erigeron : E. bonariensis L. (MH87) • E. canadensis L. (MH87) • Euthamia : E. occidentalis Nutt. (MH87) • Gutierrezia : G. californica (DC.) Torr. & A.Gray (MH87) • G. sarothrae (Pursh) Britton & Rusby (MH87) • Helenium : H. laciniatum A.Gray • Helianthus : H. annuus L. • H. petiolaris Nutt. • Heliomeris : H. multiflora Nutt. H. m. var. nevadensis (A. Nelson) W. F. Yates • Laennecia : L. coulteri (A.Gray) G.L. Nesom • Lepidospartum : L. squamatum (A.Gray) A.Gray (MH87) • Pectis • Senecio : S. flaccidus Less. • Solidago (MH87) • BORAGINACEAE Juss. Cryptantha Lehm. ex G.Don : C. intermedia (A. Gray) Greene (MH87) • Phacelia Juss. P. crenulata Torr. ex S. Watson • P. distans Benth. (MH87) • P. heterophylla Pursh (MH87) • BRASSICACEAE Burnett : Alyssum L. • Hirschfeldia : H. incana (L.) Lagr.-Foss. (MH87) • Lepidium L. L. alyssoides A.Gray • L. integrifolium Nutt. • L. thurberi Wooton • Lobularia Desv. L. maritima (L.) Desv. (MH87) • Physaria (Nutt. ex Torr. & A.Gray) A.Gray : P. gordonii (A.Gray) O’Kane & Al-Shehbaz • Sisymbrium L. S. irio L. (MH87) • CLEOMACEAE Bercht. & J. Presl : Cleome L. C. lutea Hook. • Wislizenia Engelm. W. refracta Engelm. • CONVOLVULACEAE Juss. Convolvulus L. C. arvensis L. • CRASSULACEAE J. St. -Hil. Sedum L. (MH87) • EUPHORBIACEAE Juss. Croton L. C. californicus M̹ll. Arg. (MH87) • C. setigerus Hook. (MH87) • Euphorbia L. E. albomarginata Torr. &A.Gray • FABACEAE Juss. Amorpha : A. fruticosa L. • Dalea : D. candida Willd. D. c. var. oligophylla (Torr.) Shinners • Melilotus • Prosopis : P. glandulosa Torr. P. g. var. torreyana (L.D. Benson) M.C. Johnst. • P. juliflora (Sw.) DC. • P. velutina Wooton • Psorothamnus : P. scoparius (A.Gray) Rydb. • LAMIACEAE Martinov : Mentha L. M. spicata L. (MH87) • LOASACEAE Juss. Mentzelia L. M. laevicaulis (Douglas) Torr. & A. Gray (MH87) • MALVACEAE Juss. Sphaeralcea A. St. -Hil. • PAPAVERACEAE Juss. Eschscholzia Cham. E. californica Cham. (MH87) • PLANTAGINACEAE Juss. Penstemon Schmidel : P. superbus A. Nelson • Plantago L. • POLEMONIACEAE Juss. Eriastrum Wooton & Standl. E. virgatum (Benth.) H. Mason (MH87) • POLYGONACEAE Juss. Eriogonum Michx. E. abertianum Torr. • E. fasciculatum Benth. • E. gracile Benth. (MH87) • Persicaria Mill. P. lapathifolia (L.) Delarbe (MH87) • RANUNCULACEAE Juss. Clematis L. C. ligusticifolia Nutt. • ROSACEAE Juss. Fallugia Endl. F. paradoxa (D. Don) Endl. ex Torr. • Prunus L. (MH87) • SALICACEAE Mirb. Salix L. S. exigua Nutt. (MH87) • S. lasiolepis Benth. (MH87) • S. nigra Marshall (MH87) • SAPINDACEAE Juss. Sapindus L. S. drummondii Hook. &Arn. • TAMARICACEAE Link : Tamarix L. T. ramosissima Ledeb. • VERBENACEAE J. St. -Hil. Glandularia J.F. Gmel. G. bipinnatifida (Schauer) Nutt. • Lantana L. (MH87) • ZYGOPHYLLACEAE R.Br. Larrea Cav. L. tridentata (Sessé & Moc. ex DC.) Coville.

DNA barcodes

Eight sequences available (BOLD process IDs: DIAL1098-07, DIAL1108-07, DLII755-07, DLII783- 07, DLII1085-07, BOWGF2050-10, BWTWO1316-10, BWTWO1318-10; BIN: BOLD:AAE5948). There is some shallow within-species divergence (maximum p-distance 1.33%), but this is much less than the difference between species in the L. clematisellum complex (minimum p-distance 2.78%). Lasioglossum clematisellum differs from L. clastipedion sp. nov. and L. julipile sp. nov. by 11 fixed substitutions: 18(C), 80(T), 123(C), 141(C), 286(C), 312(T), 354(T), 390(C), 426(C), 474(C), and 579(C) (Supplementary file 4). No unique fixed substitutions distinguish L. clematisellum from all other western red-tailed L. ( Dialictus ).

Remarks

Common. The male is described here for the first time. Lasioglossum clematisellum has an extremely wide distribution, but unlike the similarly-distributed L. hudsoniellum , L. clematisellum is not found at very high elevations and is more common in deserts.

Figures

Fig. 28. Lasioglossum (D.) clematisellum (Cockerell, 1904), holotype, ♀ (USNM 12064). A. Dorsal habitus. B. Lateral habitus. C. Face. D. Propodeum (not holotype). E. Metasoma. Scale bars: 1 mm.

Fig. 29. Lasioglossum (D.) clematisellum (Cockerell, 1904), ♂. A. Dorsal habitus. B. Lateral habitus. C. Face. D. Propodeum and metasoma. Scale bars: 1 mm.

Fig. 30. Georeferenced collection records of Lasioglossum (D.) clematisellum (Cockerell, 1904) (black squares) and predicted distribution by maximum entropy ecological niche modeling in Maxent (colour shading). Warmer colours indicate higher cloglog probability of occurrence.

Fig. 90. Male genitalia. A. Lasioglossum (D.) arenisaltans sp. nov. B. L. (D.) clastipedion sp. nov. C. L. (D.) clematisellum (Cockerell, 1904). D. L. (D.) hudsoniellum (Cockerell, 1919). E. L. (D.) julipile sp. nov. F. L. (D.) lilianae sp. nov. G. L. (D.) mesillense (Cockerell, 1898). H. L. (D.) meteorum sp. nov. I. L. (D.) miltolepoides sp. nov. J. L. (D.) minckleyi sp. nov. K. L. (D.) pallidellum (Ellis, 1914). L. L. (D.) pictum (Crawford, 1902). M. L. (D.) rufornatum sp. nov. N. L. (D.) spivakae sp. nov. O. L. (D.) testaceum (Robertson, 1897).P. L. (D.) vierecki (Crawford, 1904). Q. L. (D.) zephyrus (Smith, 1853). Genitalia are illustrated in dorsal view and positioned such that the gonostylus is parallel to the plane of vision. Retrorse lobes are also illustrated separately to the bottom left of each figure in lateral view, positioned parallel to the plane of vision.

Fig. 91. Tegula. A. Lasioglossum (D.) hunteri (Crawford, 1932), ♀, enlarged, reaching scutellum, with concave inner posterior margin. B. L. (D.) clematisellum (Cockerell, 1904), ♀, ovoid and not reaching scutellum. Scale bars: 1 mm.

Fig. 106. Ocellocular area. A. Lasioglossum (D.) clavicorne sp. nov., ♀, sparsely and distinctly punctate. B. L. (D.) clematisellum (Cockerell, 1904), ♀, densely punctate with punctures somewhat obscured by microsculpture. Scale bar: 0.25 mm.

Fig. 114. T1–3. A. Lasioglossum (D.) cactorum sp. nov., ♂, convex with rims strongly downcurved (arrows). B. L. (D.) clematisellum (Cockerell, 1904), ♂, flat. Scale bars: 1 mm.

Fig. 117. Mesoscutum and metasoma. A. Lasioglossum (D.) clematisellum (Cockerell, 1904), with most hair thin and simple. B. L. (D.) julipile sp. nov., with most hair densely plumose. Scale bar: 1 mm.