Tinotus trisectus Casey, 1906

Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, Zootaxa 353 (1), pp. 1-134 : 11-12

Tinotus trisectus Casey, 1906

( Figs. 2 , 7 , 23, 28–29 in Klimaszewski & Pelletier 2002)

Tinotus trisectus Casey, 1906: 321 .

Acrimea fimbriata Casey, 1911a: 15 , syn. nov.

Tinotus pallidus Casey, 1911a: 65 , syn. nov.

Tinotus brunneus Casey, 1911a: 65 .

Tinotus binarius Casey, 1911a: 66 .

Tinotus lateralis Notman, 1921: 154 .

Tinotus trisectus: Klimaszewski & Pelletier, 2002: 290 (as valid species).

Tinotus binarius: Klimaszewski & Pelletier, 2002: 290 (as synonym of T. trisectus ).

Tinotus brunneus: Klimaszewski & Pelletier, 2002: 290 (as synonym of T. trisectus ).

Tinotus lateralis: Klimaszewski & Pelletier, 2002: 290 (as synonym of T. trisectus ).

Tinotus pallidus: Klimaszewski & Pelletier, 2002: 290 (as synonym of T. caviceps Casey, 1893 ; misidentification).

Tinotus binarius: Hanley, 2002: 458 (as valid species).

Tinotus brunneus: Hanley, 2002: 458 (as valid species).

Tinotus lateralis: Hanley, 2002: 463 (as valid species).

Tinotus pallidus: Hanley, 2002: 464 (as valid species).

Tinotus trisectus: Hanley, 2002: 467 (as valid species).

(Other references are omitted)

Type material. Holotype of Tinotus trisectus : UNITED STATES: Idaho: Kootenai Co.: ♀, Coeur d'Alene, vi (H.F.Wickham) ( NMNH).

Lectotype of Acrimea fimbriata (here designated): ♂, “The Dalles, Oreg.[on (Wickham)]”, “ fimbriata Csy. ”, “TYPE USNM 39738” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype (♀; designated by Hanley (2002)) and paralectotype (♂) of Tinotus pallidus : UNITED STATES: Iowa: Linn Co.: Cedar Rapids ( NMNH).

Lectotype of Tinotus brunneus (designated by Hanley (2002)): UNITED STATES: Pennsylvania: ♂, near Philadelphia ( NMNH).

Holotype of Tinotus binarius : UNITED STATES: Arizona: Pima Co.: ♀, Tucson ( NMNH).

Additional material. UNITED STATES: California: Kern Co.: ♂, 12 mi. S Bakersfield, 20.v.1958 (E.I.Schlinger) ( CNCI) ; Tennessee: Lake Co.: ♂, Reelfoot Lk., 7 km NE Tiptonville, at black light, 8.v.1986 (J.M.Campbell) ( CNCI) .

Diagnosis. See Klimaszewski & Pelletier (2002).

Discussion. The lectotype of Acrimea fimbriata is similar to the holotype of Ti. trisectus in external characters. The shape of the aedeagus, including the structure of the internal sac, is identical in the lectotype of Acri. fimbriata and in examined males of Ti. trisectus .

Klimaszewski et al. (2002) placed Ti. pallidus in synonymy with Ti. caviceps . Unfortunately, the male paralectotype of Ti. pallidus lacks the head and pronotum which does not allow the comparison of these parts of the body with the males of Ti. caviceps which have strong medial impressions of the head disk and pronotum. However, both types of Ti. pallidus are more glossy than the types of Ti. caviceps and resemble Ti. trisectus in that respect. The aedeagus (including the structures of the internal sac) of the paralectotype of Ti. pallidus is similar to the aedeagus of Ti. trisectus , and not Ti. caviceps . This fact is clear when Figures 21 , 23 and 17–20 in Klimaszewski & Pelletier (2002) are compared. Therefore, Ti. pallidus is removed from synonymy with Ti. caviceps and placed in synonymy with Ti. trisectus Casey, 1906 .

Hanley (2002) designated the lectotype of Ti. trisectus . However, when Casey (1906) described Ti. trisectus he indicated that he had “the single type” of this species. Therefore this type is the holotype, and the lectotype designation by Hanley (2002) is redundant. Hanley (2002) also designated the lectotype of Ti. binarius . When Casey (1911a) described Ti. binarius he indicated that he had “only one example” of this species. Therefore this type is the holotype, and the lectotype designation by Hanley (2002) is redundant.

Distribution. Ti. trisectus is widespread in the United States. It is known from Oregon, California, Arizona, Idaho, Iowa, Pennsylvania and Tennessee.

Figures

FIGURES 1–4. Abdominal segment 8 of Tinotus acerbus (Casey) (lectotype of Acrimea acerba (1–2) and lectotype of Acri. resecta (3–4)). 1 — male tergum 8; 2 — male sternum 8; 3 — female tergum 8; 4 — female sternum 8. Scale bar 0.2 mm.

FIGURES 5–9. Genitalia of Tinotus acerbus (Casey) (lectotype of Acrimea acerba (5–8) and lectotype of Acri. resecta (9)). 5 — median lobe, parameral view; 6 — median lobe with details of retracted internal sac, abparameral view; 7 — median lobe, lateral view; 8 — median lobe with details of retracted internal sac, lateral view; 9 — spermatheca. Scale bar 0.1 mm (9), 0.2 mm (5– 8).

FIGURES 23–30. Genitalia of Paradilacra densissima (Bernhauer) (male, lectotype of Pa. persola Casey (23–28); female, lectotype of Pa. memnonia Casey (29); and female, lectotype of Pa. deserticola Casey (30)). 23 — median lobe, parameral view; 24 — apex of median lobe, parameral view; 25 — median lobe, lateral view; 26 — apex of median lobe, lateral view; 27 — apex of median lobe with details of retracted internal sac, parameral view; 28 — apex of median lobe with details of retracted internal sac, lateral view; 29–30 — spermatheca. Scale bar 0.1 mm (24, 26, 29– 30), 0.2 mm (23, 25, 27–28).

FIGURES 19–22. Abdominal segment 8 of Paradilacra densissima (Bernhauer) (male, lectotype of Pa. persola Casey (19–20); and female, lectotype of Pa. memnonia Casey (21–22)). 19 — male tergum 8; 20 — male sternum 8; 21 — female tergum 8; 22 — female sternum 8. Scale bar 0.2 mm.

FIGURES 15–18. Genitalia of Oxypoda amica Casey (male, lectotype of O. amica (15–17); and female from 2.5 km N Lawrence (18)). 15 — median lobe, parameral view; 16 — median lobe with details of retracted internal sac, lateral view; 17 — median lobe with details of retracted internal sac, abparameral view; 18 — spermatheca. Scale bar 0.1 mm (18), 0.2 mm (15–17).