Plagiognathus politus Uhler, 1895: 52

SCHUH, RANDALL T., 2001, Revision Of New World Plagiognathus Fieber, With Comments On The Palearctic Fauna And The Description Of A New Genus (Heteroptera: Miridae: Phylinae), Bulletin of the American Museum of Natural History 2001 (266), pp. 1-267 : 196-198

Plagiognathus politus Uhler, 1895: 52 (n. sp.). Plagiognathus politus flaveolus Knight, 1923: 434

(n. var.).

DIAGNOSIS: Recognized unequivocally by the serrate margin of the vesical flange (fig. 30); dorsum convexly rounded, pronotum declivent; and dorsum clothed with recumbent, golden, shining, simple setae (fig. 11). Occurring as distinctive early­ and late­season color morphs: early­season morph with dorsum, antennae, and all but apex of femora castaneous, cuneus with a pale lunate marking at base and posterior margin of vertex pale (fig. 11: politus 1); late­season morph with scutellum golden, often with midline of scutellum and additional areas of dorsum, including base of corium and cuneus, also golden; femora partially to largely golden (fig. 11: politus 2). Early­season morph very similar in coloration of dorsum (including lunate pale marking at base of cuneus) and femora to fuscosus (fig. 8), but antennal segment 2 largely pale in fuscosus (fig. 16) and dorsal vestiture composed of weakly flattened as well as common setae (fig. 4D), and the overall size usually smaller. Late­season morph similar in coloration to flavoscutellatus (fig. 8), but that species usually much larger and having a much narrower ves­ ical flange lacking the serrations found in politus . Also, late­season form similar in coloration to blatchleyi , but anterior pronotal lobe usually dark in politus and always pale in blatchleyi (see also Discussion under blatchleyi ).

REDESCRIPTION: Male: Relatively broadbodied, moderately large; total length 3.46– 4.12, length apex clypeus–cuneal fracture 2.41–2.75, width across pronotum 1.10– 1.33. COLORATION (fig. 11): Dorsum castaneous with a lunate pale marking at base of cuneus, corium narrowly pale adjacent to extreme base of membrane, and posterior margin of vertex pale, or scutellum and often disk of pronotum and areas of hemelytra, including pale areas mentioned above, golden; antennal segments 1 and 2 castaneous except for pale apical annulus on segment 1 (fig. 18), segments 3 and 4 infuscate; labial segments 1 and 2 castaneous, remainder of lighter coloration; venter mostly castaneous; ventral margin of pronotum, area surrounding mesothoracic spiracle, and at least part of metathoracic scent­gland evaporatory area pale; coxae, trochanters, and femora castaneous in dark colored specimens; tibiae mostly pale, spines with dark bases; tibiae dark at articulation with femora; in specimens with golden areas on dorsum, much of venter and legs golden or pale. SURFACE AND VESTITURE: Dorsum weakly granular, smooth, moderately shining. Vestiture of dorsum composed of recumbent, golden, shining, simple setae. STRUCTURE: Dorsal surface of body noticeably convexly round­ ed, hemelytra declining laterally; hemelytra broadest at point just anterior to cuneal fracture; frons nearly straight across between eyes in dorsal view, head dorsoventral in orientation, clypeus not visible from above; anteocular distance equal to diameter of antennal segment 1; head projecting below eye by 1.5 times diameter of antennal segment 1; labium reaching to about apex of hind coxae. GENITALIA (fig. 30): Body of vesica relatively stout and strongly curving, more or less U­shaped, base of vesica reaching to level of secondary gonopore; posterior apical spine elongate, nearly straight, erect relative to body of vesica, anterior spine obliquely angled relative to body of vesica, about equal in length to posterior, rather sharply bent sub­ apically; flange very broad, reaching near apex of secondary gonopore, serrate along margin at level of gonopore.

Female: Very similar to male in coloration; body not quite so elongate in appearance. Total length 3.43–3.98, length apex clypeus–cuneal fracture 2.42–2.67, width across pronotum 1.16–1.36.

HOSTS: Recorded from a variety of plants, but probably breeding primarily on the Asteraceae . Also known to feed at least in part on animal material, as recorded by Wheeler (1971).

DISTRIBUTION: Widely distributed in eastern North America, with no records from the high plains of Canada, despite extensive collecting there by Brooks and Kelton. Because of its otherwise complete absence from west of the Front Range of the Rocky Mountains, the single record from Lone Pine, California, might be considered suspect and possibly the result of mislabeling.

DISCUSSION: This is one of the most commonly encountered species of Plagiognathus in eastern North America. As noted by Knight (1941), the species routinely has two generations per year, each of which has its own distinctive coloration, the early generation being almost totally castaneous, the later being heavily marked with gold.

This species was described on the basis of a single specimen from Fort Collins, Colorado, collected on July 24 by C. F. Baker. Modern authors, including Knight, have apparently not verified whether any specimen from Uhler’s collection in the United States National Museum of Natural History actually represents the holotype and indeed I have not found any either. The concept of the species, nonetheless, has been agreed upon by most authors. I am designating a neotype (male) to ensure continued stability in the use of this name. It bears the following label data and is deposited in the United States National Museum of Natural History, Washington, D.C.

St. Anthony Pk. , Minn., August 2, 1924, H. H. Knight; H. H. Knight Collection 1976; Neotype Plagiognathus politus Uhler , det. R. T. Schuh

SPECIMENS EXAMINED: CANADA.— Nova Scotia: Kentville, July 10, 1976, L. A. Kelton, Trifolium sp. (Fabaceae) , 13 (CNC).

Ontario: Aldershot, July 6, 1955 – July 14, 1958, L. A. Kelton, 153, 9♀ (CNC). Amherstburg, July 6, 1962, Kelton and Brumpton, 43, 4♀ (CNC). Ancaster, July 1, 1954 – July 2, 1954, J. E. H. Martin, 243, 13♀ (CNC). Ancaster, July 21, 1955, L. A. Kelton, 33, 12♀ (CNC). Appin, June 19, 1962, Kelton and Brumpton, 13 (CNC). Aylmer, July 19, 1955, L. A. Kelton, Ambrosia sp. (Asteraceae) , 33, 14♀ (CNC). Bells Corners, August 7, 1962, D. Brown, Ulmus sp. (Ulmaceae) , 13 (CNC). Brantford, July 21, 1955, L. A. Kelton, 1♀ (CNC). Bryanston, June 20, 1962, Kelton and Brumpton, 13 (CNC). Burtch, July 11, 1961, L. A. Kelton, 93, 6♀ (CNC). Caledonia, July 12, 1955, L. A. Kelton, 13, 5♀ (CNC). Camden (East Ontario), June 27, 1974, D. G. Reid, 1♀ (CNC). Cayuga , July 12, 1955, L. A. Kelton, 23 (CNC). Cayuga , July 15, 1961, L. A. Kelton, 33, 4♀ (CNC). Cayuga , June 1, 1954, R. Lambert, 13 (CNC). Cayuga , June 26, 1961, Kelton and Brumpton, 1♀ (CNC). Chambers Corn, July 10, 1962, H. Blanchard, 2♀ (CNC). Chatham, August 30, 1935, W. E. Lindsay, 1♀ (CNC). Chatham, July 16, 1928 – August 3, 1928, A. B. Baird, 3♀ (CNC). Chatham, September 1, 1948, D. A. Arnott, Trifolium sp. (Fabaceae) , 33, 2♀ (CNC). Chatterton, July 24, 1956 – July 28, 1956, J. C. Martin, 13, 2♀ (CNC). Clear Creek, July 4, 1964, Kelton and Brumpton, 1♀ (CNC). Clinton, September 7, 1961, L. A. Kelton, Ambrosia sp. (Asteraceae) , 173, 9♀ (CNC). Coboura, July 11, 1981, D. J. E. Brown, 1♀ (CNC). Copenhagen, July 3, 1962, G. Thorpe, 1♀ (CNC). Copetown, July 16, 1961, Kelton and Brumpton, Picea sp. (Pinaceae) , 1♀ (CNC). Dundas, July 16, 1962, Kelton and Thorpe, 33, 4♀ (CNC). Dunnville, August 3, 1957, R. Lambert, 13 (CNC). Dunnville, July 9, 1962, G. Thorpe, 13, 2♀ (CNC). Eden, July 2, 1962, H. Blanchard, 1♀ (CNC). Edwards, Patterson’s Farm, June 23, 1991, M. D. Schwartz, 43, 9♀ (CNC). Effingham, August 2, 1961, L. A. Kelton, 1♀ (CNC). Effingham, July 12, 1955, L. A. Kelton, 53, 21♀ (CNC). Eramosa, September 5, 1961, Kelton and Brumpton, 13, 7♀ (CNC). Fonthill, September 8, 1947, J. H. H. Phillips, Prunus sp. (Rosaceae) , 1♀ (CNC). Footes Bay, July 25, 1962, Kelton and Thorpe, Spiraea sp. (Ro­ saceae), 1♀ (CNC). Fort Erie, July 10, 1955, L. A. Kelton, 23, 3♀ (CNC). Fuller, August 1, 1972, D. G. Reid, Solidago canadensis (Asteraceae) , 1♀ (CNC). Fuller, July 2, 1971 – July 28, 1971, D. G. Reid, Solidago sp. (Asteraceae) , 83, 2♀ (CNC). Fuller, July 8, 1967, Solidago sp. (Asteraceae) , 1♀ (CNC). Goderich, July 2, 1962, G. Thorpe, Anethum (Apiaceae) , 1♀ (CNC). Goderich, July 2, 1962, H. Blanchard, 23, 1♀ (CNC). Grand Bend, September 6, 1954, C. D. F. Miller, 33 (CNC). Grassie, June 21, 1962, Kelton and Thorpe, 13 (CNC). Haggersville, August 23, 1961, Kelton and Brumpton, 1♀ (CNC). Haggersville, July 9, 1962, Kelton and Thorpe, Ambrosia sp. (Asteraceae) , 83, 15♀ (CNC). Hamilton, July 16, 1967, L. A. Kelton, 1♀ (CNC). Hamilton, June 27, 1968, L. A. Kelton, Ambrosia sp. (Asteraceae) , 133, 16♀ (CNC). Hamilton, July 6, 1955, L. A. Kelton, 53, 3♀ (CNC). Harrow, August 30, 1961, J. Brumpton, 13 (CNC). Ipperwash, July 11, 1962, Kelton and Thorpe, 1♀ (CNC). Jordan, July 17, 1961, L. A. Kelton, Crataegus sp. (Rosaceae) , 113, 6♀ (CNC). Jordan, June 7, 1962, Kelton and Thorpe, 13 (CNC). Kinburn, July 18, 1957, J. E. H. Martin, 13 (CNC). Kincardine, September 7, 1961, Kelton and Brompton, 23, 1♀ (CNC). Kingsville, July 17, 1955, L. A. Kelton, 113, 10♀ (CNC). Kingsville, July 7, 1962, G. Thorpe, 13, 2♀ (CNC). Kingsville, June 18, 1962 – June 19, 1962, Kelton and Thorpe, Crataegus sp. (Rosaceae) , 143, 5♀ (CNC). Kintore, July 10, 1962, Kelton and Thorpe, Picea sp. (Pinaceae) , 23, 1♀ (CNC). Leamington, June 26, 1931 – July 2, 1931, G. S. Walley, 153, 9♀ (CNC). Leamington, June 29, 1961, G. Brumpton, 13 (CNC). Leamington, September 12, 1961 – September 13, 1961, Kelton and Brumpton, Ambrosia sp. (Asteraceae) , 73, 10♀ (CNC). Marmora, April 4, 1952 – July 16, 1952, E. H. N. Smith, Medicago sativa (Fabaceae) , 2♀ (CNC). Marmora, July 3, 1952 – July 18, 1952, J. R. McGillis, 23, 4♀ (CNC). Marmora, July 5, 1952 – July 24, 1952, J. R. Vockeroth, 43, 1♀ (CNC). Marmora, July 8, 1952 – July 14, 1952, C. Boyle, 33, 3♀ (CNC). Marmora, September 10, 1952, J. F. McAlpine, Rhus aromatica (Anacardiaceae) , 13 (CNC). McGregor, July 7, 1962, G. Thorpe, 13 (CNC). Montecello Hill, June 28, 1968, L. A. Kelton, 3♀ (CNC). Mt. Pleasant, August 22, 1961, L. A. Kelton, 83, 8♀ (CNC). Mt. Pleasant, July 10, 1958, L. A. Kelton, Salix sp. (Salicaceae) , 103, 19♀ (CNC). Mt. Vernon, July 10, 1962, Kelton and Thorpe, 23 (CNC). Nepean, Piney Forest, July 18, 1991 – August 28, 1991, M. D. Schwartz, 23, 7♀ (CNC). Newry, July 12, 1962, Kelton and thorpe, Salix sp. (Salicaceae) , 13, 2♀ (CNC). Niagara Falls, August 10, 1961, L. A. Kelton, Erigeron sp. (Asteraceae) , 33, 8♀ (CNC). Niagara Falls, July 7, 1955, L. A. Kelton, 53, 6♀ (CNC). Norway Point, Lake of Bays, July 26, 1919, J. Mc­ Dunnough, 13 (CNC). Norwich, July 19, 1962, Kelton and Thorpe, 1♀ (CNC). Oakland, July 17, 1962, Kelton and Thorpe, 1♀ (CNC). Ojibway, August 30, 1961, J. Brumpton, 53, 2♀ (CNC). Ottawa, July 11, 1957, J. E. H. Martin, 23, 2♀ (CNC). Ottawa, June 28, 1957 – September 22, 1952, E. H. N. Smith, Medicago sp. (Fabaceae) , 33 (CNC). Pelee Island, July 3, 1931, G. S. Walley, 233, 14♀ (CNC). Pelee Island, September 12, 1961, Kelton and Brumpton, 153, 19♀ (CNC). Port Burwell, July 3, 1962, G. Thorpe, 33, 3♀ (CNC). Port Burwell, July 3, 1962, H. Blanchard, 63, 4♀ (CNC). Port Rowan, July 12, 1962, H. Blanchard, 23, 1♀ (CNC). Port Rowan, July 13, 1955, L. A. Kelton, 13, 1♀ (CNC). Port Rowan, July 4, 1962, L. A. Kelton, 1♀ (CNC). Pt. Anne, April 9, 1948, J. C. Martin, 13 (CNC). Pt. Pelee, July 6, 1962, Kelton and Thorpe, 13 (CNC). Pt. Pelee, July 9, 1931, G. S. Walley, 43, 2♀ (CNC). Pt. Pelee, June 28, 1961, Kelton and Brumpton, 13 (CNC). Pt. Pelee, September 11, 1961, L. A. Kelton, 13 (CNC). Pt. Pelee, September 9, 1954, R. M. Mason, 1♀ (CNC). Queenston, July 23, 1963, L. A. Kelton, 23, 7♀ (CNC). Queenston, July 8, 1955, L. A. Kelton, 33, 5♀ (CNC). Rockaway, June 21, 1962, Kelton and Thorpe, 73, 2♀ (CNC). Selkirk, July 9, 1962, Kelton and Brumpton, Abies sp. ( Pinaceae ), 23, 3♀ (CNC). Selkirk, July 9, 1962, Kelton and Brumpton, Nepeta sp. (Lamiaceae) , 13, 5♀ (CNC). Selkirk, July 9, 1962, Kelton and Brumpton, Rubus sp. (Rosaceae) , 23, 3♀ (CNC). Shipka, July 10, 1962, Kelton and Thorpe, Ulmus americana (Ulmaceae) , 13, 9♀ (CNC). Simcoe, July 13, 1955, L. A. Kelton, 13, 1♀ (CNC). Simcoe, July 2, 1915, 1♀ (CNC). Simcoe, July 20, 1915, H. G. Crawford, 1♀ (CNC). Smithville, June 10, 1962, Kelton and Brumpton, Rubus sp. (Rosaceae) , 1♀ (CNC). South Gloucester, Blossom Park, Greenbelt, June 9, 1991, M. D. Schwartz, 1♀ (CNC). Spencerville, August 31, 1939 – September 9, 1939, G. H. Hammond, 3♀ (CNC). St. Catharines, August 8, 1961 – September 9, 1961, L. A. Kelton, 243, 12♀ (CNC). St. Catharines, July 18, 1963, L. A. Kelton, 3♀ (CNC). St. Catharines, July 9, 1955, L. A. Kelton, 93, 23♀ (CNC). St. Lawrence Islands Natl. Park, Aubrey Island, September 15, 1976, W. Reid, 23 (CNC). St. Lawrence Islands Natl. Park, Thwartway Island, July 25, 1976, W. Reid, 13 (CNC). St. Thomas, July 19, 1955, L. A. Kelton, 23 (CNC). Stanford, July 12, 1961, L. A. Kelton, 113, 14♀ (CNC). Stirling, June 26, 1962, G. Thorpe, 23, 3♀ (CNC). Stittsville, August 9, 1962, D. Brown, Salix sp. (Salicaceae) , 1♀ (CNC). Strathroy, July 19, 1925 – July 20, 1925, H. F. Hudson, 13, 3♀ (CNC). Thornhill, August 15, 1961, L. A. Kelton, 13, 8♀ (CNC). Tillsonburg, July 14, 1955 – July 11, 1958, L. A. Kelton, Artemisia sp. (Asteraceae) , 153, 25♀ (CNC). Tillsonburg, July 3, 1962, H. Blanchard, 23, 4♀ (CNC). Tillsonburg, June 18, 1962 – June 20, 1962, Kelton and Thorpe, 23, 2♀ (CNC). Trenton, June 27, 1911, Evans, 13 (CNC). Trenton, September 11, 1910, Evans, 13 (CNC). Turkey Point, July 12, 1962, H. Blanchard, 1♀ (CNC). Vienna, July 15, 1961, L. A. Kelton, 23 (CNC). Vienna, July 18, 1962, Kelton and Thorpe, Salix sp. (Salicaceae) , 13, 3♀ (CNC). Vineland, July 11, 1955 – July 22, 1963, L. A. Kelton, 103, 7♀ (CNC). Virgil, August 8, 1961, L. A. Kelton, 33, 17♀ (CNC). Virgil, July 9, 1955, L. A. Kelton, 33, 7♀ (CNC). Vittoria, July 13, 1962, H. Blanchard, Rubus sp. (Rosaceae) , 13 (CNC). Vittoria, July 14, 1962, G. Brumpton, Salix sp. (Salicaceae) , 1♀ (CNC). Windsor, July 7, 1962, G. Thorpe, Quercus sp. (Fagaceae) , 1♀ (CNC). Woodslee, August 31, 1961, G. Brumpton, 1♀ (CNC). Woodslee, July 7, 1962, G. Thorpe, Platanus sp. (Platanaceae) , 13, 1♀ (CNC). Quebec: Fabre, July 5, 1963, L. A. Kelton, 13 (CNC). Gatineau National Park, North End, June 25, 1981, D. J. E. Brown, 1♀ (CNC). Hemmingford, August 29, 1916, J. I. Beaulne, 23, 5♀ (CNC). Hull, June 16, 1955, L. A. Kelton, 13 (CNC). Knowlton, June 26, 1929 – August 12, 1929, G. S. Walley, 13, 2♀ (CNC). La Trappe, August 28, 1933, J. Ouellet, 13, 1♀ (TAMU). Montreal, Beaulieu, August 15, 1905, 1♀ (CNC). Quyon, July 22, 1953, L. A. Kelton, 23 (CNC). USA.— Alabama: Garland Co.: near Fountain Lake on Rt 7, June 11, 1987, T. J. Henry and A. G Wheeler, Jr., Salix sp. (Salicaceae) , 13 (USNM). Arkansas: Marion Co.: Yellville, September 11, 1926, H. H. Knight, 33 (USNM). Pulaski Co.: Little Rock, May 5, 1943, Stahevitch, 13 (LACM). Sebastian Co.: Huntington, June 15, 1948, B.T. McDermott, 13 (KU). California: Inyo Co. : Lone Pine, July 28, 1940, D. E. Hardy, 13 (KU). Colorado: Arapohoe Co.: Highline Canal at Belleview Ave., June 25, 1981, D. A. Polhemus, 13 (JTP). Denver Co.: Denver, June 15, 1916, E. C. Jackson, 13 (USNM). El Paso Co.: Colorado Springs, 5915 ft, E. S. Tucker, 13 (KU). Weld Co.: No specific locality, July 2, 1978, G. E. Bohart, 23 (UCB). Connecticut: 5.5 mi N of Stamford, July 15, 1970, W. Ford, 13 (AMNH). Cheshire, September 16, 1966, L. and C. W. O’Brien, 33 (UCB). Colebrook, Colebrook Resevoir, August 21, 1970, F. P. Maroney, 23 (AMNH). Eagleville, June 22, 1977, D. Laston, 13 (AMNH). East Hartford, Tunxis State Forest, July 20, 1970, F. P. Maroney, 13 (AMNH). East Suffield, Connecticut River, July 21, 1970, F. P. Maroney, 23 (AMNH). Eastford, July 17, 1976, J. A. Slater, 13 (AMNH). Hall Meadow State Park, Torrington, August 4, 1970, F. P. Maroney, 13 (AMNH). Hartford Res., West Hartford, July 23, 1970, F. P. Maroney, 43 (AMNH). New Haven, August 23, 1934, M. E. Griffith, 13 (KU). New Haven, July 20, 1964, B. H. Walden, 13 (CAS). New Haven, July 6, 1914, M. P. Zappe, 13 (CAS). New Haven, September 4, 1911, C. E. Olsen, 73 (CAS). Pachaug State Forest, August 24, 1964, J. A. Slater, 13 (AMNH). Poquonock, Farmington River, July 22, 1970, F. P. Maroney, 343 (AMNH). Rocky Hill, August 18, 1970, F. P. Maroney, 23 (AMNH). Somers, LaChance Farm, July 8, 1965, 13 (AMNH). Sound Beach, July 23, 1910, 13 (AMNH). Storrs, July 12, 1955, R. M. Baranowski, 33 (AMNH). Storrs, July 23, 1954, R. M. Baranowski, 43 (AMNH). Storrs, July 6, 1964, 33 (AMNH). Storrs, June 28, 1977, D. Leston, 33 (AMNH). Trumbull, Easton Reservoir, September 8, 1970, F. P. Maroney, 13 (AMNH). Waterbury, September 13, 1969, C. W. O’Brien, 13 (UCB). West Hartford, M. D. C. Lab, July 17, 1970, W. Ford, 13 (AMNH). Windsor, July 6, 1965, D. E. Leonard, 53 (AMNH). Georgia: Towns Co.: Hiawassee, August 19, 1957, L. A. Kelton, 13 (CNC). Illinois: Urbana, August 25, 1930, H. H. Knight, 23 (AMNH). Cumberland Co.: Greenup, August 20, 1962, L. D. Anderson, 13 (UCR). Henderson Co.: Oquawka, June 13, 1932, H. L. Dozier, 13 (AMNH). La Salle Co.: Ottawa, June 1, 1942, F. C. Werner, 23 (AMNH). Piatt Co.: Monticello, August 16, 1932, T. H. Frison, 13 (AMNH). Pope Co.: Herod, June 24, 1932, Ross, Dozier, and Park, 13 (AMNH). Pulaski Co.: Mounds, September 18, 1931, C. W. Johnson , 13 (AMNH). Randolf Co.: Chester, June 30, 1939, D. E. Hardy, 13 (KU). Scott Co.: Bluffs, August 29, 1951, A. T. McClay, 303 (UCD). Warren Co.: Roseville, September 1, 1943, R. I. Sailer, 13 (USNM). Indiana: Kosciusko Co.: 2 mi N of Warsaw, September 1, 1966, C. W. O’Brien, 13 (UCB). Marion Co.: No specific locality, October 6, 1928, W. S. B., 73 (UCB). Tippecanoe Co.: Lafayette, August 14, 1969, L. B. O’Brien, 13 (UCB). Wayne Co.: 14 km W of Richmond, July 4, 1966, Paul H. Arnaud, Jr., 13 (CAS). Iowa: Adair Co.: Adair, July 25, 1960, P. M. Marsh, 13 (UCD). Boone Co.: Ledges State Park, July 22, 1928, G. S. Walley, 2♀ (CNC). Ledges State Park, June 26, 1953, J. C. Schaffner, 1♀ (TAMU). No specific locality, June 22, 1962, J. C. Schaffner, Asclepias syriaca (Asclepiadaceae) , 73, 3♀ (TAMU). Pilot Mound, August 30, 1960, J. C. Schaffner, 33, 3♀ (TAMU). Clinton Co.: Clinton, June 23, 1928, G. S. Walley, 23 (CNC). DeWitt, June 21, 1928, G. S. Walley, 43 (CNC). Dickinson Co.: 5 mi W of Milford, June 18, 1963, J. C. Schaffner, 53, 3♀ (TAMU). Cayler Prairie, June 11, 1963 – June 30, 1963, J. C. Schaffner, Salix sp. (Salicaceae) , 33, 1♀ (TAMU). Iowa Lakeside Lab., June 15, 1963 – June 27, 1963, J. C. Schaffner, Fraxinus pennsylvanica (Oleaceae) , 33, 1♀ (TAMU). Iowa Lakeside Lab., June 15, 1963 – June 27, 1963, J. C. Schaffner, Quercus macrocarpa (Fagaceae) , 33, 2♀ (TAMU). Henry Co.: 3 mi NW of Salem, T70N, R7W, Sec. 9, June 16, 1962, J. C. Schaffner, Erigeron sp. (Asteraceae) , 13, 1♀ (TAMU). 5 mi SW of Mt. Pleasant, June 30, 1976 – July 1, 1976, J. C. Schaffner, 63, 12♀ (TAMU). 6 mi SE of Mt. Pleasant, T71N R7W Sec. 28, June 16, 1962, J. C. Schaffner, Melilotus alba (Fabaceae) , 43, 2♀ (TAMU). 7 mi SW of Mt. Pleasant, July 4, 1964, J. C. Schaffner, 13, 2♀ (TAMU). 7 mi W of Mt. Pleasant, July 2, 1976, J. C. Schaffner, 33, 7♀ (TAMU). Mt. Pleasant, September 19, 1932, C. Hall, 1♀ (TAMU). Mt. Pleasant, September 27, 1928, Bruhn, 1♀ (TAMU). Olds, August 27, 1963, S. G. Wellso, 1♀ (TAMU). Johnson Co. : 2 mi W of Solon, August 29, 1966, C. W. O’Brien, 13 (UCB). Iowa City, July 25, 1971, L. A. Kelton, Robinia sp. (Fabaceae) , 3♀ (CNC). Muscatine Co.: Wilton, July 8, 1927, Harris and Johnston, 2♀ (TAMU). Polk Co.: Mitchellville, Thomas Mitchell Co. Park, June 29, 1980, K. and R. Schmidt, 663 (AMNH). Scott Co.: Pleasant Valley, July 5, 1928, G. S. Walley, 203, 11♀ (CNC). Story Co.: Ames, August 23, 1952, J. C. Schaffner, 13, 1♀ (TAMU). Ames, August 28, 1953 – September 2, 1953, J. C. Schaffner, 83, 10♀ (TAMU). Ames, July 7, 1930, H. G. Johnston, 1♀ (TAMU). Ames, June 11, 1962, J. C. Schaffner, Heracleum maximum (Apiaceae) , 43, 4♀ (TAMU). Ames, June 21, 1929, H. M. Harris, 13 (TAMU). Ames, June 21, 1971 – September 25, 1967, H. H. Knight, 223, 38♀ (USNM). Ames, June 25, 1927 – July 5, 1927, H. G. Johnston, 133, 5♀ (TAMU). Ames, June 25, 1962, J. C. Schaffner, Ambrosia trifida (Asteraceae) , 23, 1♀ (TAMU). Ames, Pleat Bog Valley, July 22, 1928, G. S. Walley, 63, 6♀ (CNC). Ames, September 24, 1949, W. Downes, 13 (OSU). Ames, September 25, 1927, H. M. Harris, 2♀ (TAMU). Ames, September 30, 1928, H. M. Harris, 2♀ (TAMU). Ames, September 9, 1931, H. M. Harris, 13, 2♀ (TAMU). Van Buren Co.: Farmington, July 14, 1927, Harris and Johnston, 13 (CNC). Warren Co.: 1.5 mi E of Hartford, July 5, 1976 – July 6, 1976, J. C. Schaffner, 213, 21♀ (TAMU). Kansas: Atchison Co.: No specific locality, July 17, 1924, R. H. Beamer, 113 (KU). Butler Co.: Leon, June 20, 1940, L. J. Lipovski, 103 (KU). Decatur Co.: No specific locality, July 15, 1925, R. H. Beamer, 13 (KU). Doniphan Co.: No specific locality, August 22, 1921, V. J. Brown, 13 (KU). No specific locality, July 20, 1924, R. H. Beamer, 23 (KU). Douglas Co.: Lawrence vicinity, September 1, 1969, V. P. Gapud, 43 (KU). Lawrence, September 21, 1951, A. A. Hubert, 13 (KU). Lawrence, September 4, 1941, R. H. Beamer, 13 (KU). No specific locality, 900 ft, 333 (KU). No specific locality, July 9, 1924, P. B. Lawson, 53 (KU). No specific locality, June 14, 1928, P. B. Lawson, 13 (KU). No specific locality, June 26, 1975, T. W. Oldham, 33 (TAMU). No specific locality, October 18, 1944, R. H. Beamer, 23 (KU). Sibleyville, September 30, 1952, A. A. Hubert, 13 (KU). Leavenworth Co.: No specific locality, June 24, 1924, E. P. Breakey, 13 (KU). Linn Co.: No specific locality, 862 ft, R. H. Beamer, 13 (KU). Mc Pherson Co.: No specific locality, June 28, 1923, R. H. Beamer, 13 (KU). Miami Co.: No specific locality, R. H. Beamer, 103 (KU). Montgomery Co.: Coffeyville, June 24, 1939, L. W. Hepner, 23 (KU). No specific locality, 798 ft, R. H. Beamer, 13 (KU). Morris Co. : No specific locality, July 31, 1923, R. H. Beamer, 13 (KU). Republic Co.: No specific locality, July 10, 1925, Howard Deny, 23 (KU). Rice Co.: No specific locality, July 3, 1923, L. C. Woodruff, 23 (KU). Saline Co.: No specific locality, July 13, 1923, R. H. Beamer, 13 (KU). Wyandotte Co.: No specific locality, June 22, 1924, R. H. Beamer, 13 (KU). Maine: Lincoln Co.: Whitefield, N. Banks, 23 (AMNH). Penobscot Co.: Orono, July 14, 1905, 13 (CAS). Maryland: Cecil Co.: Conowingo, September 18, 1950, W. F. Chamberlain, 13 (TAMU). Frederick Co.: Middletown, June 13, 1985, A. G. Wheeler, Jr., 13 (PDA). Lanham Co.: Lanham, June 25, 1967, Paul Oman, 33 (OSU). Montgomery Co.: Great Falls, July 4, 1963, D. C. and K. A. Rentz, 13 (CAS). Plummers Island, August 31, 1989, W. E. Steiner, 13 (USNM). Silver Spring, June 25, 1967, Paul Oman, 23 (OSU). Prince Georges Co.: College Park, September 21, 1966, L. and C. W. O’Brien, 23 (UCB). Unknown Co.: Wolfville, June 26, 1914, 13 (USNM). Washington Co.: Clear Spring Exit on I­70, June 19, 1979, A. G. Wheeler, Jr., Achillea millefol­ ium ( Asteraceae ), 1♀ (TAMU). Hagerstown, August 23, 1915, H. L. Parker, 33, 5♀ (USNM). Hagerstown, July 7, 1916, H. L. Parker, 13 (CAS). Massachusetts: Barnstable Co.: Woods Hole, C. W. Johnson , 13 (AMNH). Essex Co.: Beach Bluff, July 10, 1915, H. M. Parshley, 63 (CAS). Beach Bluff, June 22, 1914, H. M. Parshley, 283 (CAS). Marblehead, July 24, 1914, H. M. Parshley, 13 (CAS). Saugus, September 2, 1915, H. M. Parshley, 13 (CAS). Saugus, September 5, 1914, H. M. Parshley, 43 (CAS). Swampscott, August 19, 1914, H. M. Parshley, 13 (CAS). Swampscott, July 13, 1915, H. M. Parshley, 33 (CAS). Franklin Co.: Northfield, August 10, 1898, F. Bowditch, 13 (AMNH). Sunderland, Mt. Toby, September 23, 1917, H. M. Parshley, 33 (CAS). Hampshire Co.: Northampton, August 13, 1918, H. M. Parshley, 13 (CAS). Middlesex Co.: Auburndale, C. W. Johnson , 33 (AMNH). Framingham, August 15, 1914, C. A. Frost, 13 (CAS). Holliston, N. Banks, 333 (AMNH). Lexington, N. Banks, 73 (AMNH). Sherborn, August 14, 1915, C. A. Frost, 13 (CAS). Sherborn, September 7, 1919, A. P. Morse, 123 (AMNH). Norfolk Co.: Brookline, June 11, 1905, H. M. Parshley, 13 (CAS). Cohasset, September 21, 1914, H. M. Parshley, 33 (CAS). Dedham, C. W. Johnson , 23 (AMNH). Needham, September 3, 1911, 23 (CAS). Wellesley, July 11, 1909, E. P. Van Duzee, 13 (CAS). Wellesley, September 6, 1900, 13 (CAS). Suffolk Co.: Boston, June 17, 1914, H. M. Parshley, 13 (CAS). Forest Hills, September 19, 1914, H. M. Parshley, 13 (CAS). Forest Hills, September 30, 1915, H. M. Parshley, 23 (CAS). Worcester Co.: Petersham, 13 (AMNH). Michigan: Emmet Co.: No specific locality, August 10, 1950, Doris Gardner, 13 (OSU). Ingham Co.: East Lansing, September 14, 1965, E. D. Evans, 33 (AMNH). Mason, September 12, 1958, H. D. Niemczyk, 23, 2♀ (USNM). Van Buren Co.: Lawton, Huzzy’s Lake, July 22, 1984, J. A. Jackman, 1♀ (TAMU). Minnesota: Ramsey Co.: 4 mi W of St. Anthony Park, June 30, 1964, J. C. Schaffner, 1♀ (TAMU). St. Anthony Park, August 2, 1924, H. H. Knight, 13, 1♀ (TAMU). St. Anthony Park, August 2, 1924, H. H. Knight, 233, 54♀ (USNM). St. Paul, September 3, 1923, H. H. Knight, Ambrosia trifida (Asteraceae) , 53, 2♀ (USNM). Mississippi: Adams Co.: Natchez, May 15, 1931, H. G. Johnston, 43, 6♀ (TAMU). Natchez, May 15, 1931, H. G. Johnston, Aster sp. (Asteraceae) , 43, 7♀ (TAMU). Lowndes Co.: Columbus, June 20, 1929, H. G. Johnston, 13, 1♀ (TAMU). Oktibbeha Co.: Starkville, June 13, 1929, H. G. Johnston, 13, 1♀ (TAMU). Tallahatchie Co.: Charleston, August 31, 1925, H. M. Harris, 1♀ (TAMU). Missouri: Boone Co.: Columbia , September 24, 1966, F. D. Parker, 13 (UCD). Rt 1, 0.6 mi S of jct of Hwy N, September 2, 1982, R. L. Blinn, Ambrosia trifida (Asteraceae) , 73, 3♀ (USNM). Franklin Co.: Meramec State Park, June 30, 1968, L. A. Kelton, 13 (CNC). Jackson Co.: Atherton, June 13, 1915, C. F. Adams, 63 (CAS). Kansas City, F. Rogers, 23 (KU). Kansas City, Willow Creek, I­435 and Wormill Rd., June 11, 1977, H. N. Greenbaum, 13 (TAMU). Jasper Co.: Joplin, June 30, 1968, L. A. Kelton, 33, 6♀ (CNC). Vernon Co.: 4 mi W of Montevallo, June 24, 1966, J. C. Schaffner, 13 (TAMU). Wayne Co.: Williamsville, July 7, 1955, E. C. Becker, 1♀ (CNC). Montana: Toole Co.: Shelby City Campground, July 7, 1972, G. C. Gaumer, 1♀ (TAMU). Nebraska: Cass Co.: Rest Area on I­ 80, 2 mi W of Platte Rd., June 29, 1980, K. Schmidt, 23 (AMNH). Hall Co.: Grand Island, July 8, 1964, H. H. Knight, 2♀ (USNM). Nance Co.: 14 mi SE of Fullerton, August 27, 1966, L. B. O’Brien, 13 (UCB). Nemaha Co.: Peru, Peru State College, August 22, 1975, A. G. Wheeler, Jr., Ambrosia trifida (Asteraceae) , 13, 7♀ (PDA). Red Willow Co.: Indianola, A. P. Morse, 23 (AMNH). New Hampshire: Carroll Co.: Notchland, August 20, 1934, M. E. Griffith, 13 (KU). Coos Co.: Bretton Woods, August 31, 1934, J. D. Beamer, 13 (KU). Mount Washington, A. T. Slosson, 23 (AMNH). Rockingham Co.: Hampton, October 22, 1922, S. Albert Shaw, 13 (CAS). New Jersey: Bergen Co.: Closter, June 26, 1962, S. J. Hessel and J. A. Woods, 13 (AMNH). Closter, September 17, 1972, J. G. Rozen, 13 (AMNH). Mahwah, July 3, 1962, Rozen, Statham, Woods, and Hessel, 13 (AMNH). Ramsey, August 23, 1921, 13 (AMNH). Ramsey, July 20, 1917, 23 (AMNH). Ramsey, July 6, 1912, 13 (AMNH). Ramsey, September 3, 1917, 23 (AMNH). Ramsey, September 7, 1918, 13 (AMNH). Rutherford, E. L. Dickerson, 13 (AMNH). Sicomac, September 8, 1922, T. M. Schott, 23 (AMNH). Mercer Co.: Mercerville, August 24, 1910, 13 (AMNH). Princeton, September 18, 1966, C. W. O’Brien, 23 (UCB). Middlesex Co.: 2 mi SW of New Brunswick, September 8, 1966, C. W. O’Brien, 23 (UCB). New Brunswick, 83 (AMNH). New Brunswick, September 25, 1943, 13 (AMNH). Passaic Co.: Bearfort Mountains, September 30, 1965, P. and B. Wygodzinsky and J. A. Woods, 13 (AMNH). Paterson, 13 (AMNH). Somerset Co.: Somerville, July 12, 1922, Schott, 13 (AMNH). Sussex Co.: Baleville, June 21, 1981, Robert Schmidt, ex Asteraceae , 23, 2♀ (AMNH). Unknown Co.: Milltown, 13 (AMNH). New Mexico: Lincoln Co.: White Mountains, 8200 ft, Ruidoso Natl. Forest, 13 (USU). Otero Co.: 3 mi E of Cloudcroft, July 19, 1976, John D. Pinto, 153 (UCR). New York: Albany Co.: Rensselaerville, Huyck Preserve, June 29, 1977, R. T. Schuh, 13 (AMNH). Bronx Co.: Mosholu, July 12, 1919, 13 (AMNH). Mosholu, July 9, 1891, 103 (AMNH). Mosholu, September 9, 1910, 233 (AMNH). Van Cortlandt Park, Bronx, July 4, 1962, S. J. Hessel, 23 (AMNH). Cattaraugus Co.: Gowanda, August 29, 1907, E. P. Van Duzee, 13 (CAS). Cayuga Co. : Aurora, July 21, 1909, V. L. Zabriski, 23 (AMNH). Chautauqua Co.: Fredonia, July 21, 1946, L. D. Beamer, 13 (KU). Coumbia Co.: Chatham, August 30, 1904, A. P. Morse, 383 (AMNH). Erie Co.: Buffalo, August 24, 1898, E. P. Van Duzee, 23 (CAS). Elma, August 25, 1912, E. P. Van Duzee, 13 (CAS). Hamburg, July 1, 1906, E. P. Van Duzee, 13 (CAS). Hamburg, July 5, 1904, E. P. Van Duzee, 43 (CAS). Hamburg, July 9, 1905, E. P. Van Duzee, 13 (CAS). Hamburg, June 21, 1908, E. P. Van Duzee, 43 (CAS). Hamburg, June 26, 1898, E. P. Van Duzee, 83 (CAS). Hamburg, June 28, 1896, E. P. Van Duzee, 23 (CAS). Hamburg, September 6, 1909, E. P. Van Duzee, 43 (CAS). Holland, July 22, 1946, R. H. Beamer, 13 (KU). Lancaster, July 25, 1946, L. D. Beamer, 23 (KU). Essex Co.: Whiteface Mountain, 4600–4872 ft, July 19, 1962, J. R. Vockeroth, 13 (CNC). Genesee Co.: Batavia, July 5, 1914, H. H. Knight, 13 (CNC). Batavia, September 2, 1915, H. H. Knight, 13 (CAS). Kings Co.: Flatbush, July 2, 1895, V. L. Zabriski, 23 (AMNH). Flatbush, Long Island, June 26, 1893, V. L. Zabriski, 13 (AMNH). Flatbush, Long Island, September 5, 1892, 13 (AMNH). Flatbush, September 14, 1897, V. L. Zabriski, 33 (AMNH). Parkville, Long Island, May 30, 1891, Beutenmulier, 23 (AMNH). Water Works, Flatbush, Long Island, July 10, 1893, V. L. Zabriski, 13 (AMNH). Water Works, Flatbush, Long Island, September 18, 1892, V. L. Zabriski, 33 (AMNH). Nassau Co.: Old Brookville, 140 ft, August 21, 1973, M. Schwartz, 13 (AMNH). Sea Cliff, Long Island, N. Banks, 13 (AMNH). Sea Cliff, Long Island, N. Banks, 43 (AMNH). New York Co.: Inwood Park, New York City, July 19, 1970, T. Gidaspow, 13 (AMNH). Oneida Co. : near Sangerfield on Rt 20, September 6, 1975, A. G. Wheeler, Jr., Solidago sp. (Asteraceae) , 13 (PDA). Orange Co.: Fort Montgomery, September 16, 1923, F. M. Schott, 13 (AMNH). Goshen, September 7, 1910, 23 (AMNH). Pine Island, September 8, 1910, 203 (AMNH). Pine Island, September 9, 1914, 33 (AMNH). Orleans Co.: No specific locality, June 27, 1928, V. A. Little, 13 (TAMU). Rockland Co.: 5 mi NW of Tuxedo Park , July 3, 1962, Rozen, Statham, Woods, and Hessel, 13 (AMNH). Suffern, September 17, 1910, 43 (AMNH). Suffolk Co.: Amagansett, Long Island, September 15, 1910, William T. Davis, 13 (AMNH). Cold Spring Harbor, Long Island, August 5, 1920, Parshley, 13 (CAS). Cold Spring Harbor, Long Island, July 4, 1914, H. M. Parshley, 23 (CAS). Greenport, Long Island, September 18, 1946, 43, 4♀ (USNM). Tompkins Co.: Ithaca, Cornell Plantations off Judd Falls Road, May 26, 1974, T. J. Henry and A. G. Wheeler, Jr., Cornus stolonifera (Cornaceae) , 13 (PDA). Ithaca, July 27, 1920, H. H. Knight, 73, 3♀ (USNM). Ithaca, N. Banks, 13 (AMNH). Ithaca, September 4, 1973, M. Schwartz, 13 (AMNH). Ludlowville, September 1, 1980, L. L. Pechuman, Bidens sp. (Asteraceae) , 43, 4♀ (PDA). Ring Wood, July 25, 1928, V. A. Little, 23 (TAMU). Unknown Co.: Big Island, September 8, 1910, 93 (AMNH). Taugannock Falls, August 10, 1928, V. A. Little, 1♀ (TAMU). Warren Co.: Hague, July 17, 1915, 33 (AMNH). Lake George, August 21, 1893, V. L. Zabriski, 123 (AMNH). Lake George, September 3, 1908, V. L. Zabriski, 13 (AMNH). Westchester Co.: Bryn Mawr Park, September 16, 1910, 23 (AMNH). Chappaqua, June 29, 1919, 13 (AMNH). White Plains, June 28, 1919, J. R. T. B., 13 (CAS). Yonkers, July 9, 1891, Beutenmuller, 23 (AMNH). North Carolina: Highlands Co.: Whiteface Cove near Highlands, August 17, 1957, L. A. Kelton, 23, 3♀ (CNC). Jackson Co.: Tuckasegee, August 31, 1957, L. A. Kelton, 1♀ (CNC). Macon Co.: Bridal Veil Falls, June 20, 1967, H. Greenbaum, 13 (KU). Franklin, 2000 ft, June 17, 1957, J. R. Vockeroth, 23 (CNC). Franklin, August 16, 1957, L. A. Kelton, 3♀ (CNC). Highlands, August 11, 1957 – August 29, 1957, L. A. Kelton, 523, 48♀ (CNC). Highlands, August 21, 1957 – August 29, 1957, W. R. Richards, 123, 5♀ (CNC). Madison Co.: Hot Springs, A. T. Slosson, 13 (AMNH). Mecklenburg Co.: Near Matthews, Rt 51 1 mi W of Rt 16, September 3, 1973, A. G. Wheeler, Jr., Oenothera biennis (Onagraceae) , 1♀ (TAMU). Swain Co.: Great Smoky Mountains, Willets, 2000 ft, August 24, 1930, N. Banks, 13 (AMNH). Unknown Co.: Red Hill, August 14, 1957, L. A. Kelton, Salix sp. (Salicaceae) , 13 (CNC). Watauga Co.: Blowing Rock to Linville, 3000–4000 ft, September 8, 1930, N. Banks, 13 (AMNH). Yancey Co.: Black Mountains, 53 (AMNH). Black Mountains, summit, July 25, 1906, W. Beutenmuller, 13 (AMNH). Valley of Black Mountains, July 5, 1906, W. Beutenmuller, 183 (AMNH). North Dakota: Traill Co.: No specific locality, August 14, 1923, A. A. Nichol., 13 (USNM). No specific locality, August 19, 1923, A. A. Nichol, 73, 4♀ (USNM). Ohio: Franklin Co.: Columbus, Mock Park, July 14, 1984, S. M. Clark, 13 (TAMU). Worthington, July 15, 1951, R. C. Ballard, 13 (PUC). Licking Co.: Blackhand Gorge State Nature Preserve, July 14, 1988, P. W. Kovarik, 13 (TAMU). Lorain Co.: Amherst, July 1, 1933, H. J. Reinhard, 33, 2♀ (TAMU). Sandusky Co.: Rt 81 off Rt 51, 3 mi W of Green Springs, July 14, 1974, P. S. B., 23 (CAS). Stark Co.: Massillon, August 21, 1962, Lawrence K. Ertle, 13 (USU). Summit Co.: Akron, June 23, 1987, P. W. Kovarik, 1♀ (TAMU). Barberton, August 23, 1986, L. J. Lipovski, 523 (KU). Barberton, June 22, 1937, L. J. Lipovski, 13 (KU). No specific locality, June 22, 1936, Louis J. Lipovski, 23 (KU). No specific locality, L. J. Lipovsky, 113 (KU). Pennsylvania: Allegheny Co.: Pittsburgh, July 5, 1915, S. L. Mason, 23 (CAS). Bedford Co.: Breezewood, July 15, 1983, R. S. Miller, 13 (TAMU). E of Bedford on Rt 30, August 30, 1979, A. G. Wheeler, Jr., Eupatorium fistulosum (Asteraceae) , 1♀ (PDA). Berks Co.: Gibralter, September 7, 1971, A. G. Wheeler, Jr., Coronilla varia (Fabaceae) , 13 (PDA). Leesport, July 1, 1970, 73, 6♀ (PDA). N of Moselem on Rt 666, June 11, 1974, A. Gnagey, 13 (PDA). Virginville, June 6, 1968, P. Vaurie, 13 (AMNH). Blair Co.: Altoona Cemetery, July 7, 1976, T. J. Henry, Rhus glabra (Anacardiaceae) , 13, 2♀ (PDA). Altoona on Rt 764, June 27, 1974, A. G. Wheeler, Jr. et al., Cornus sp. (Cornaceae) , 43, 3♀ (PDA). Near Martinsburg, George Bridenbough Nursery, June 27, 1974, A. G. Wheeler, Jr., Elaeagnus angustifolia (Elaeagnaceae) , 3♀ (PDA). Williamsburg, J. B. Farm, July 17, 1973, B. Stinner, Corylus americana (Betulaceae) , 13, 2♀ (PDA). Bucks Co.: Horseshoe Bend, Neshaminy Creek NE of Jamison, July 4, 1965, Wilton Ivie, 13 (AMNH). Jamison, June 26, 1968, Wilton Ivie, 13 (AMNH). Butler Co.: Butler, Eisler Nurseries, July 9, 1976, R. Henry, Tamarix sp. (Tamaricaceae) , 4♀ (PDA). Centre Co.: Bellefonte, September 12, 1973, A. G. Wheeler, Jr., Quercus palustris (Fagaceae) , 2♀ (PDA). Spring Mills, Penn Nursery, September 14, 1971, D. Stehr, Robinia pseudoacacia (Fabaceae) , 1♀ (PDA). State College, Rt 322, July 6, 1976, T. J. Henry, Rhus glabra (Anacardiaceae) , 3♀ (PDA). State College, University Drive, September 12, 1973, A. G. Wheeler, Jr., 13, 1♀ (PDA). Woodward, July 2, 1971, A. G. Wheeler, Jr., 1♀ (PDA). Chester Co.: Marshalton, Fethcroft Nursery, June 11, 1974, A. G. Wheeler, Jr., Tanacetum vulgare (Asteraceae) , 33, 4♀ (PDA). Columbia Co. : Bloomsburg, September 30, 1914, 13 (AMNH). Cumberland Co.: Hagerstown, September 5, 1973, D. Stehr, Abutilon theoprasti (Malvaceae) , 13, 6♀ (PDA). Newville, Conifer Hill Nursery, July 12, 1972, Pinus strobus (Pinaceae) , 23, 2♀ (PDA). Pennsylvania Turnpike, 9 mi E of Rt 997, July 2, 1980, A. G. Wheeler, Jr., 13 (PDA). Dauphin Co.: 7 mi E of Harrisburg, July 17, 1971, E. E. Simons, 13 (PDA). 7 mi N of Harrisburg, August 16, 1973, T. J. Henry and A. G. Wheeler, Jr., Vernonia sp. (Asteraceae) , 13 (PDA). Conewago Township, Cedar Road, May 11, 1974, A. G. Wheeler, Jr., Robinia pseudoacacia (Fabaceae) , 13 (PDA). Devonshire Heights, August 29, 1958, 1♀ (PDA). East Hanover Township, Crooked Hill Road, May 22, 1974, T. J. Henry and A. G. Wheeler, Jr., Ulmus occidentalis (Ulmaceae) , 13 (PDA). Harrisburg, Agriculture Building, June 19, 1974, T. J. Henry, Sambucus canadensis (Caprifoliaceae) , 13 (PDA). Harrisburg, Cameron Street, June 20, 1974, B. Stimmel, Sambucus canadensis (Caprifoliaceae) , 23 (PDA). Harrisburg, Farmshow Road, June 29, 1971, D. Tritt, 13 (PDA). Harrisburg, June 29, 1900, W. Reinick, 23 (PDA). Harrisburg, Wildwood Park, July 19, 1900, W. Reinick, 13 (PDA). Hummelstown, June 29, 1919, 1♀ (PDA). Lower Paxton Township, June 2, 1975, A. G. Wheeler, Jr., Rhodotypos scandens (Rosaceae) , 13 (PDA). Nyes and Willoughby Roads, June 16, 1975, T. J. Henry and A. G. Wheeler, Jr., Vitis sp. (Vitaceae) , 13 (PDA). Delaware Co.: No specific locality, W.J. Gerhard, 43 (CAS). Erie Co.: Fairview on Rt 98, August 22, 1973, T. J. Henry and A. G. Wheeler, Jr., Helianthus sp. (Asteraceae) , 13 (PDA). Fairview, Fairview Nurseries, August 22, 1973, T. J. Henry and A. G. Wheeler, Jr., Ambrosia sp. (Asteraceae) , 13 (PDA). Presque Isle, July 21, 1920, E. M. Craighead, 13 (PDA). Fayette Co.: Royal, A&J Nursery, July 8, 1973, D. Trelka, Pinus sylvestris (Pinaceae) , 1♀ (PDA). Franklin Co.: Chambersburg, August 17, 1921, J. R. Stehr, 13, 4♀ (PDA). Marion, July 22, 1922 – August 17, 1922, J. R. Stehr, 2♀ (PDA). Quincy, August 20, 1971, R. Colburn, 23 (PDA). Lancaster Co.: Elizabethtown, July 1, 1974, A. G. Wheeler, Jr., Tilia cordata (Tiliaceae) , 13 (PDA). Near Millersville, Herr Nursery, August 15, 1974, T. J. Henry, Abelia sp. (Caprifoliaceae) , 13 (PDA). Rt 441, Falmouth Access Area, August 16, 1979, A. G. Wheeler, Jr., Cassia fasiculata (Fabaceae) , 13 (PDA). Lebanon Co.: I­80 E of Grantville exit, June 20, 1974, T. J. Henry and A. G. Wheeler, Jr, Sambucus canadensis (Caprifoliaceae) , 13 (PDA). Monroe Co.: Delaware Water Gap, H. G. Barber, 13, 1♀ (USNM). Northampton Co.: Easton, June 28, 1954, J. W. Green, 63 (CAS). Easton, September 15, 1956, J. W. Green, 263 (CAS). Wind Gap, July 9, 1954, J. W. Green, 73 (CAS). Northumberland Co.: 2 mi NE of Sunbury, June 18, 1975, A. G. Wheeler, Jr., Kalmia latifolia (Lamiaceae) , 33 (PDA). Perry Co.: Amity Hall, May 31, 1979, A. G. Wheeler, Jr., Rosa multiflora (Rosaceae) , 13 (PDA). Near Livermore on Rt 15, June 5, 1974, A. G. Wheeler, Jr., Verbascum thapsi (Scrophulariaceae) , 13 (PDA). Sullivan Co.: 4 mi N of LaPorte on Rt 220, August 8, 1974, A. G. Wheeler, Jr., Spiraea latifolia (Rosaceae) , 13 (PDA). Unknown Co.: Marysville, August 25, 1900, 1♀ (PDA). Marysville, August 25, 1900, W. Reinick, 23 (PDA). Venago Co.: Sugar Creek, August 21, 1973, T. J. Henry and A. G. Wheeler, Jr., Oenothera sp. (Onagraceae) , 3♀ (PDA). Washington Co.: Taylorstown exit off Hwy 70, July 3, 1966, Paul H. Arnaud Jr., 273 (CAS). Wayne Co.: Honesdale, October 3, 1913, C. E. Olsen, 13 (AMNH). Near Calicoon, New York, Curtis Nursery, June 18, 1974, A. G. Wheeler, Jr., Gleditsia triacanthos (Fabaceae) , 13, 3♀ (PDA). Westmoreland Co.: Charter Oak, July 11, 1917, J. B. Kirk, 1♀ (PDA). York Co.: Dillsburg, Cooke Farm, September 15, 1971, D. Stehr, Coronilla varia (Fabaceae) , 13 (PDA). Manchester, Dauber’s Farm, July 3, 1973, Lehnian and Null, 13 (PDA). York, Walker Nursery, June 19, 1975, K. R. Valley, Spiraea sp. (Rosaceae) , 13, 2♀ (PDA). Rhode Island: Unknown Co.: No specific locality, Parshley, 13 (CAS). Washington Co.: Kingston, 13 (CAS). South Carolina: Greenville Co.: Greenville, July 14, 1976, R. S. Peigler, 13 (TAMU). Greenville, June 7, 1984, R. S. Peigler, 1♀ (TAMU). Greenville, May 27, 1977, R. S. Peigler, 13 (TAMU). Taylors, May 28, 1977, R. S. Peigler, 23 (TAMU). Tigerville, August 20, 1930, R.H. Beamer, 13 (KU). South Dakota: Brookings Co.: Brookings, June 17, 1921, H. C. Severin, 13 (CAS). Roberts Co.: Lake Traverse, 12 mi SE of Sisseton, July 19, 1974, Blair Tollefson, 13 (UCB). Tennessee: Montgomery Co.: Clarksville, July 4, 1939, J. D. Beamer, 33 (KU). Unknown Co.: 6 mi W of North Carolina border, 1200 ft, September 15, 1930, N. Banks, 23 (AMNH). Texas: Bandera Co. : Lost Maples State National Area, May 25, 1985, P. W. Kovarik and R. W. Jones, 13 (TAMU). Brazos Co.: Bryan, May 6, 1965 – May 10, 1975, J. C. Schaffner, 593, 65♀ (TAMU). College Station, May 10, 1933, H. G. Johnston, 13 (TAMU). College Station, May 6, 1975, J. C. Schaffner, 2♀ (TAMU). College Station, October 1, 1927, H. G. Johnston, 1♀ (TAMU). Little Brazos River, Hwy 21, May 20, 1970, V. V. Board, 23 (TAMU). Burleson Co.: No specific locality, May 11, 1964, Jackson, 13, 1♀ (TAMU). Somerville Lake, April 29, 1969, V. V. Board, 13 (TAMU). DeKalb Co.: 6 mi SE of Jefferson, May 31, 1998, W. F. Chamberlain, 13 (TAMU). Gonzales Co.: 0.5 mi E of Harwood, May 13, 1969, V. V. Board, 23, 1♀ (TAMU). Palmetto State Park, April 19, 1969, V. V. Board, 13 (TAMU). Palmetto State Park, April 22, 1970 – May 4, 1970, V. V. Board, 83, 3♀ (TAMU). Palmetto State Park, May 4, 1970, Board and Schaffner, 183, 6♀ (TAMU). Kerr Co.: Kerrville, June 2, 1997, G. M. Chamberlain, 1♀ (TAMU). Kerrville, June 20, 1996, W. F. Chamberlain, 3♀ (TAMU). Kerrville, May 11, 1992, W. F. Chamberlain, 1♀ (TAMU). Kerrville, May 12, 1994, G. M. Chamberlain, 13 (TAMU). Kerrville, May 18, 1993, W. F. Chamberlain, 23, 1♀ (TAMU). Kerrville, May 3, 1998 – May 11, 1998, G. M. Chamberlain, 23, 1♀ (TAMU). Kerrville, May 31, 1997 – June 1, 1997, W. F. Chamberlain, 13, 1♀ (TAMU). Milam Co.: Gause 3 mi N at Sugar Loaf Mt., May 27, 1995, W. Godwin, H. Howden, A. Howden, 1♀ (TAMU). Real Co.: 4 mi S of Garven Store, April 9, 1995, W. F. Chamberlain, 13 (TAMU). Travis Co.: Shelberg Tract near Cypress Creek arm of Lake Travis, May 8, 1994, J. C. Schaffner, 33, 12♀ (TAMU). Vicinity of Long Hollow Creek, May 7, 1994, J. C. Schaffner, 93, 1♀ (TAMU). Vicinity of Long Hollow Creek, May 7, 1994, M. Quinn, E. Riley, R. Wharton, Quercus buckleyi (Fagaceae) , 1♀ (TAMU). Uvalde Co.: Garner State Park, May 5, 1961, 1♀ (TAMU). Wood Co.: 6 mi NW of Hawkinson, Highway 14, May 23, 1998, W. Godwin, 13, 1♀ (TAMU). Vermont: Grand Isle Co.: South Hero, A. P. Morse, 23 (AMNH). Orange Co.: Bradford, July 23, 1973, J. Amaral, 23, 1♀ (TAMU). Windham Co.: Brattleboro, June 15, 1908, H. M. Parshley, 13 (CAS). Windsor Co.: Springfield, August 22, 1934, R. H. Beamer, 13 (KU). Woodstock, A. P. Morse, 13 (AMNH). Virginia: Arlington Co.: Glencarlyn, N. Banks, 33 (AMNH). Clarke Co.: 3 mi E of Berryville, Shenandoah River, July 24, 1977, N. L. Herman, 13 (AMNH). Fairfax Co.: Great Falls, N. Banks, 13 (AMNH). Falls Church Co.: Falls Church, N. Banks, 213 (AMNH). Giles Co.: Stinking Creek, 1850 ft, June 25, 1969, George W. Byers, 13 (KU). Patrick Co.: Woolwine, September 1, 1946, R. H. Beamer, 13 (KU). Washington, D. C.: June 25, 1905, E. P. Van Duzee, 13 (CAS). West Virginia: Jefferson Co.: No specific locality, August 22, 1980, T. L. Mason, 63, 9♀ (PDA). Mineral Co.: Patterson Creek Road N of Grant County line, June 10, 1979, A. G. Wheeler, Jr., Ulmus americana (Ulmaceae) , 13 (PDA). Nicholas Co.: Craigsville, June 24, 1978, A. G. Wheeler, Jr., Picea abies (Pinaceae) , 1♀ (TAMU). Pocahontus Co.: N of Slatyfork on Rt 219, August 15, 1982, Verbesina alternifolia (Asteraceae) , 2♀ (TAMU). Preston Co.: Cathedral State Park, July 29, 1989, C. L. Staines, 23, 2♀ (USNM). Randolf Co.: No specific locality, August 8, 1975, L. Butler, 1♀ (TAMU). Unknown Co.: Aurora, August 21, 1904, O. Heidemann, 13 (CU). Wood Co.: No specific locality, July 25, 1971 – July 27, 1971, L. H. Rolston, 3♀ (TAMU). Wisconsin: Milwaukee Co.: Milwaukee, August 22, 1960, E. P. Brakey, 33 (KU).