Macrotylus verticalis Uhler, 1894: 272

SCHUH, RANDALL T., 2001, Revision Of New World Plagiognathus Fieber, With Comments On The Palearctic Fauna And The Description Of A New Genus (Heteroptera: Miridae: Phylinae), Bulletin of the American Museum of Natural History 2001 (266), pp. 1-267 : 237-239

Macrotylus verticalis Uhler, 1894: 272 (n. sp.). Plagiognathus fusciflavus Knight, 1929c: 267 (n.

sp.). NEW SYNONYMY.

DIAGNOSIS: Recognized by medium to moderately large body size, the uneven brownish coloration of the dorsum with some lighter, diffuse, mostly longitudinal markings (fig. 14), the tumid frons with dark transverse striae, the brown calli, and the black, suberect, bristlelike setae on the pronotum and costal margin of the hemelytra. Also body appearing slender and transversely convexly rounded as viewed from above. Possibly confused with lineatus (fig. 9) because of the longitudinal markings, and also with moerens (fig. 10) on the basis of bristlelike setae, distribution, and occurrence on ruderal vegetation, but distinguished because both of those species with frons entirely dark and lacking obvious striate markings, lineatus lacking black bristlelike setae on dorsum, and moerens with black setae over entire dorsum and with distinctive genitalia.

REDESCRIPTION: Male: Elongate, mediumsized to moderately large; total length 3.90– 4.99, length apex clypeus–cuneal fracture 2.65–3.30, width across pronotum 1.06–1.24. COLORATION (fig. 14): General coloration uneven brown; vertex and frons mostly pale, frons with brown, transverse striae; calli darker than remainder of pronotum; hemelytra with a diffuse longitudinal pattern of brown and pale; membrane fumose, veins pale; antennae black except for pale apical annulus on segment 1 (fig. 19); labium and entire venter brown; femora with numerous dark spots on dirty yellow background; dorsal tibial spines with dark spots at bases; tibiae dark at femoral articulation. SURFACE AND VESTITURE: Dorsum weakly granular, smooth, dull to very weakly shining. Dorsum clothed with recumbent, pale to golden, simple setae, much of pronotum and costal margin of hemelytra with suberect, black, bristlelike setae. STRUCTURE: Body elongate, relatively slender, nearly parallelsided, with dorsum transversely convexly rounded, body appearing somewhat cylindrical; frons bulging as viewed from above, clypeus mostly visible; anteocular distance 1.3 times diameter of antennal segment 1; head projecting below level of eye by diameter of antennal segment 1; labium relatively long, reaching just past apex of hind coxae. GENITALIA (fig. 33): Vesica more or less J­shaped, body relatively slender, base falling just below base of secondary gonopore; posterior apical spine long, slender, erect relative to body of vesica, anterior spine slightly longer, slender, smoothly curving, and weakly angled relative to body of vesica; flange extending past body, terminating at base of secondary gonopore.

Female: Shorter and more strongly ovoid than male; coloration similar to male. Total length 3.64–4.23, length apex clypeus–cuneal fracture 2.49–2.96, width across pronotum 1.07–1.34.

HOSTS: Recorded as occurring on a variety of herbaceous, mostly annual, plant species.

DISTRIBUTION: Western North America, ranging from British Columbia and Idaho in the north, south to Baja California. The record from British Columbia (Bowser) is from Schwartz and Scudder ( 2000) as fusciflavus Knight ; I did not examine their specimens.

DISCUSSION: This is one of the most commonly collected species of Plagiognathus in California, and as the locality data indicate, it has been collected at only a limited number of localities outside the state. It is obviously most frequently collected by sweeping roadside or other ruderal vegetation, an observation that would explain why it is common in collections.

My identification of this species is based on three males and one female deposited in the California Academy of Sciences bearing the locality label ‘‘S. Francisco, Cal.’’ They are also labeled as ‘‘paratype’’ (probably by Van Duzee, judging from the yellow­orange labels) and as ‘‘Uhler type’’. Two apparent discrepancies exist with regard to the labeling of these specimens and their candidacy as Uhler types. First, Uhler indicated that ‘‘Several specimens are in the collection, labelled ‘Cal. 2.’ ’’ He went on to say that varieties occurred in San Diego and other places in California and that he had seen specimens from southern Baja California. It is therefore not clear whether he actually examined the specimens which were at some point labelled as ‘‘Uhler type’’. Obviously, also, if these specimens were examined by Uhler, they could not be paratypes.

Van Duzee (1914) had some doubts about the identity of verticalis when he commented that Uhler’s description was ‘‘very inadequate’’. Later, Van Duzee (1916b) placed verticalis in the orthotyline genus Pseudopsallus ; he noted that the types were from California.

Nothing appears to have been written on verticalis since the works of Uhler and Van Duzee. I have examined most, if not all, of the phyline Miridae from the collections of the California Academy of Sciences and from the National Museum of Natural History, with the latter institution being the repository of the Uhler collection. I have not encountered any specimens that bear the label ‘‘Cal. 2’’.

Plagiognathus verticalis , as here conceived, is distinct from all other North American Plagiognathus species and conforms in most respects to the way most previous authors have used the name. I have not desig­ nated a lectotype, on the chance that additional searching may produce a different conclusion concerning this taxon and whether the above­mentioned specimens represent material examined by Uhler.

Knight (1929c) described the species Plagiognathus fusciflavus from LaGrande, in northeastern Oregon. Examination of the holotype male and paratype female indicates that this species is the same as the taxon I am calling verticalis Uhler , corresponding closely in the linear pattern of the markings on the hemelytra, the dark clypeus and maxillary plate, the dark calli, and the pale veins of the membrane; both specimens examined by Knight are badly rubbed and devoid of setae on the dorsum. Furthermore, the type locality for fusciflavus is within the range of specimens I have otherwise identified as verticalis .

SPECIMENS EXAMINED: MEXICO.— Baja California Norte: 14.2 mi S of Tecate, May 29, 1980, Brown and Faulkner, 3♀ (SDNH). 18.2 mi E of San Telmo, May 23, 1980, Brown and Faulkner, 23, 7♀ (SDNH). 22 km W of Parque Sierra San Pedro Martir, 1150 m, April 25, 1985, R. T. Schuh and B. M. Massie, Adenostoma fasciculatum (Rosaceae) , 13 (AMNH). 38 km E of Rt 1 toward Parque San Pedro Martir, 400 m, April 24, 1985, R. T. Schuh and B. M. Massie, Viguiera laciniata (Asteraceae) , 143, 12♀ (AMNH). 5.2 mi E of Tecate, May 28, 1981, Brown and Faulkner, 13, 1♀ (SDNH). 7 km W of Parque Sierra San Pedro Martir, 1720 m, April 25, 1985, R. T. Schuh and B. M. Massie, Ceanothus cuneatus (Rhamnaceae) , 23, 8♀ (AMNH). Ensenada, May 9, 1936, C. E. Norland, 13, 1♀ (LACM). San Quentin, May 13, 1938, C. E. Norland, 13, 3♀ (LACM). Santa Marta , May 23, 1980, Brown and Faulkner, 2♀ (SDNH). Santo Domingo, May 12, 1938, C. E. Norland, 13 (LACM). Tecate , 3.4 mi S of El Condor, May 15, 1982, M. D. Schwartz, Lupinus sp. (Fabaceae) , 83, 3♀ (AMNH). USA.— California: Alameda Co. : 12 mi S of Livermore, Arroyo Mocho , June 7, 1969, P. Opler, ex Apiaceae , 83, 6♀ (UCB). Amador Co.: Fiddletown, May 29, 1960, W. E. Simonds, 13 (CAFA). Plymouth, May 29, 1960, W. E. Simonds, Calochortus sp. (Liliaceae) , 43 (CAFA). Butte Co. : Pentz, May 22, 1928, H. H. Keifer, 4♀ (CAS). Calaveras Co. : 4 mi S of Railway Flat, 2800 ft, May 18, 1969, E. G. Linsley, 13 (UCB). 4.8 km S of West Point, July 21, 1980, Stanley C. Williams, 1♀ (CAS). Angels Camp, May 22, 1930, E. P. Van Duzee, 23 (CAS). Mokelumne Hill, May 18, 1931, R. L. Usinger, 283, 47♀ (UCB). Del Norte Co. : Little Grayback, 4600 ft, July 24, 1969, J. Powell, 13, 2♀ (UCB). El Dorado Co. : 2 mi N of Kelsey, June 23, 1967, W. J. Turner, 83, 4♀ (UCB). 2 mi N of Placerville, June 7, 1962, John T. Doyen, 1♀ (UCB). 3 mi S of Camino, July 3, 1948, P. D. Hurd, 1♀ (UCB). 3 mi W of Placerville, June 5, 1962, J. T. Doyen, 2♀ (UCB). Camino , June 30, 1949, W. F. Chamberlain, 1♀ (TAMU). Chile Bar, July 5, 1948, R. A. Smith, 1♀ (UCB). Greenwood, June 21, 1967, P. F. Warner, 43, 4♀ (UCB). Riverton, June 5, 1968, S. Paul Welles, 73, 3♀ (UCB). Snowline Camp, July 30, 1948, P. D. Hurd, 13, 3♀ (UCB). Snowline Camp, June 30, 1948, W. F. Chamberlain, 13 (TAMU). Fresno Co. : 5 mi W of Avery, May 17, 1990, R. S. Miller, 13, 3♀ (TAMU). Dunlap, May 29, 1949, W. Dwight Pierce, Delphinium sp. (Ranunculaceae) , 23, 3♀ (LACM). San Benito, Big Panoche Creek , April 21, 1967, J. Powell, 13, 2♀ (UCB). Humboldt Co. : Dinsmores, June 18, 1939, Brunson P. Bliven, 103, 20♀ (CAS). Dyerville, July 28, 1963, 13 (CAS). Laribee Valley , July 3, 1938, Brunson P. Bliven, 1♀ (CAS). Kern Co. : Cerro Noroeste, SW Kern County , 8200 ft, July 15, 1965, J. Powell, 13 (UCB). Lake Co. : 2 mi S of Middletown, April 4, 1963, W. Turner, 1♀ (UCB). 5 mi NW of Middletown, June 4, 1963, P. F. Warner, 2♀ (UCB). Blue Lake , June 3, 1954, S. M. Fidal, 1♀ (UCD). E of Clear Lake on State Hwy, June 16, 1936, Brunson P. Bliven, 23 (CAS). Lake Pillsbury , June 8, 1977, W. W. Middlekauff, 23, 1♀ (UCB). Middle Creek , June 7, 1932, R. L. Usinger, Eriodictyon californicum (Hydrophyllaceae) , 33, 2♀ (UCB). N fork Cache Creek on Highway 20, May 17, 1961, D. R. Miller, 13, 1♀ (UCD). Lassen Co. : Susanville, June 20, 1975, T. R. Haig, 13, 2♀ (CAFA). Los Angeles Co. : 22 mi S of Palmdale, 3000 ft, May 30, 1981, J. T. Polhemus, 43, 9♀ (JTP). 5 mi N of Escondido Beach, 1500 ft, March 16, 1967, J. Powell, Gilia achillaefolia (Polemoniaceae) , 13 (UCB). Beverly Glen, June 13, 1957, R. X. Schick, 13 (LACM). Claremont, C. F. Baker, 13 (CAS). Claremont, May 15, 1954, J. D. Hall, 23, 4♀ (UCD). Griffith Park Bird Sanctuary, L. J. Muchmore, 1♀ (LACM). Lancaster, May 19, 1937, E. P. Van Duzee, 2♀ (CAS). Mint Canyon, May 25, 1937, E. P. Van Duzee, 1♀ (CAS). Mint Canyon, Oaks, April 20, 1932, E. P. Van Duzee, 1♀ (CAS). No specific locality, April 1, 1900, Koebele Collection, 23, 1♀ (CAS). Pasadena, May 25, 1909, Grinell, 23, 9♀ (CAS). Pebbly Beach Canyon , Santa Catalina Island, March 31, 1968, J. Powell, 1♀ (UCB). Santa Monica Mountains , 23, 1♀ (LACM). Santa Monica Mountains , May 15, 1966, B. Stabile, 2♀ (LACM). Tanbark Flat, June 17, 1956 – June 25, 1952, A. S. Menke, B. Tinglof, 33, 1♀ (LACM). Tanbark Flat, June 18, 1956, C. L. Whey, 23, 2♀ (UCB). Tanbark Flat, June 24, 1952, J. K. Hester, 33, 1♀ (UCD). Tapia Park, April 9, 1966, 13 (LACM). Temescal Canyon, April 12, 1979, 1♀ (LACM). Madera Co. : 4 mi W of Bass Lake, 3000 ft, July 1, 1946, H. P. Chandler, 13 (UCB). Biledo Meadows, July 27, 1946, T. O. Thatcher, 1♀ (UCB). Coarsegold, 2000 ft, June 29, 1946, H. P. Chandler, 13 (UCB). Marin Co. : Pt. Reyes, June 16, 1935, E. P. Van Duzee, 1♀ (CAS). Mariposa Co. : 1.9 mi W of Mt. Bullion, May 30, 1959, G. I. Stage, 23 (UCB). 3 mi SW of Miami Ranger Station, 4000 ft, June 29, 1946, H. P. Chandler, 43, 1♀ (CAS). 6 mi SW of Mt. Bullion, May 30, 1959, G. I. Stage, 4♀ (CAS). Mendocino Co. : 10 mi W of Covelo, June 28, 1961, D. C. Rentz, 23 (CAS). 2 mi S of Hopland, June 19, 1958, T. R. Haig, 1♀ (UCD). 4 mi S of Hopland, May 9, 1961, R. L. Langston, 1♀ (UCB). 4 mi W of Eel River R. S., Mendocino National Forest , June 12, 1972, J. Doyen, 163, 13♀ (UCB). 7 mi W of Eel River R. S., Mendocino National Forest , 1450 ft, June 10, 1972, J. Powell, 63, 2♀ (UCB). Eel River R. S., Mendocino National Forest , June 13, 1972, J. Powell, Aesculus californica (Hippocastanaceae) , 4♀ (UCB). Hopland , May 9, 1926, E. P. Van Duzee, 2♀ (USNM). Hopland , May 9, 1926, E. P. Van Duzee, 13, 6♀ (CAS). Navarro, June 1, 1967, W. W. Middlekauff and D. C. Rentz, 13, 2♀ (UCB). Ukiah, June 8, 1931, R. L. Usinger, Verbena sp. (Verbenaceae) , 63, 9♀ (UCB). Modoc Co. : 2.5 mi S of Rt 139 toward Lookout, 1440 m, July 6, 1979, R. T. and Joe Schuh, Lupinus sp. (Fabaceae) , 33, 2♀ (AMNH). 21.6 mi NW of Canby, 1440 m, July 1, 1979, R. T. Schuh and B. M. Massie, Wyethia helianthoides (Asteraceae) , 133, 23♀ (AMNH). 24.7 mi NW of Canby, 1375 m, July 1, 1979, R. T. Schuh and B. M. Massie, 2♀ (AMNH). 8 mi S of Rt 139 toward Lookout, 1420 m, July 6, 1979, R. T. and Joe Schuh, Sidalcea sp. (Malvaceae) , 63, 5♀ (AMNH). Adin Pass, July 8, 1977, T. R. Haig, 1♀ (CAFA). Fandango Pass Summit, Warner Mts. , 1890 m, July 3, 1979, R. T. Schuh and B. M. Massie, 13, 1♀ (AMNH). Monterey Co. : 6 mi W of Greenfield, Wiley Ranch , 1200 ft, May 3, 1975, 23, 1♀ (UCB). Bradley, April 23, 1917, E. P. Van Duzee, 13 (CAS). Bryson, April 26, 1917, E.P. Van Duzee, 73 (CAS). Bryson, May 18, 1920, E. P. Van Duzee, 83, 1♀ (CAS). Paloma Creek , 3 air mi NE of Arroyo Seco Guard Station, 900 ft, May 5, 1975, W. W. Middlekauff, 13 (UCB). Paraiso Springs, June 2, 1932, L. S. Stein, 1♀ (CAS). Pleyto, May 21, 1920, E. P. Van Duzee, 1♀ (CAS). Napa Co. : 5 mi E of Conn Dam, June 5, 1964, J. Powell, Eriophyllum lanatum (Asteraceae) , 13 (UCB). Samuel Spring, May 18, 1955, E. I. Schlinger, 13, 2♀ (UCD). Nevada Co. : 1 mi W of Hobart Mills, Prosser Creek , 5800 ft, June 17, 1966, W. Gagne, 1♀ (UCB). 2 mi N of Cisco, July 27, 1962, J. T. Doyen, 13 (UCB). 8 mi S of Grass Valley, May 18, 1920, E. P. Van Duzee, 13 (CAS). Nevada City , June 17, 1936, Brunson P. Bliven, 13, 4♀ (CAS). Sagehen Creek , July 5, 1953, K. R. Daily, 33, 1♀ (UCD). Orange Co. : Lower Santa Ana Canyon, Green River Camp , May 9, 1933, E. P. Van Duzee, 63, 3♀ (CAS). Placer Co. : Squaw Valley , July 15, 1961, E. Ball, 2♀ (CAS). Plumas Co. : Butte Lake , July 5, 1932, R. L. Usinger, 23, 4♀ (UCB). Caribou, July 6, 1932, R. L. Usinger, 53, 10♀ (UCB). Clio, July 8, 1952, M. Cazier and R. Schrammel, 1♀ (AMNH). Frenchman Reservoir, 5000 ft, June 18, 1966, W. Gagne, 1♀ (UCB). Keddie, June 20, 1941, Fred H. Rindge, 13, 1♀ (UCB). Riverside Co. : 10.5 mi SE of Santa Margarita , May 16, 1980, J. D. Pinto, 2♀ (UCR). 5 mi S of Sage, April 15, 1965, C. A. Toschi, 13 (UCB). 8 km N of Mareno Bad Lands, 1200 ft, May 19, 1981, N. J. Gunther, 1♀ (UCR). 8 mi SE of Hemet, Bautista Canyon , May 1, 1974, J. D. Pinto, 13, 1♀ (UCR). Cactus Valley , T 65 R1 E Sec 8, April 29, 1980, J. N. Chandler, 23, 1♀ (UCR). Deep Canyon, September 23, 1963, E. Schlinger and M. Irwin, 13 (UCR). Menifee Valley , hills on W end, 1800 ft, April 26, 1977 – May 14, 1978, J. D. Pinto, Salvia mellifera (Lamiaceae) , 143, 6♀ (UCR). Menifee Valley , hills on W end, 1800 ft, May 11, 1978, J. D. Pinto and R. T. Schuh, 43, 8♀ (AMNH). Milepost 23.75 on Rt 60, April 21, 1980, Russell and Schwartz, 13 (AMNH). Quail Valley , Coastal Sage Scrub Community, May 19, 1977, J. D. Pinto, Brassica sp. (Brassicaceae) , 13, 6♀ (AMNH). Riverside, May 8, 1962, Joyce Kleinjan, 23, 1♀ (UCR). San Jacinto Mountains , 5000 ft, June 7, 1908, F. Grinnell, Jr., 3♀ (CAS). San Jacinto Mountains , April 26, 1953, A. Fukushima, 13 (LACM). San Jacinto Mountains , Hemet Reservoir, May 23, 1940, R. L. Usinger, Lupinus sp. (Fabaceae) , 13, 2♀ (UCB). San Jacinto Mountains , Pinon Flat, May 31, 1940, R. Husbands, 13 (UCB). Vandevanter Flat, San Jacinto Mountains , June 4, 1945, C. D. Michener, 13 (UCB). Sacramento Co. : Carmichael, May 26, 1946, 13 (SDNH). Folsom, May 30, 1933, H. H. Keifer, Wyethia sp. (Asteraceae) , 13 (CAFA). Orangevale, May 25, 1930, H. H. Kelfer, Lupinus sp. (Fabaceae) , 53, 4♀ (LACM). San Benito Co. : 3.8 mi on New Idria Rd. from Panoche Rd. (17.9 mi. off I­5), May 21, 1978, F. C. Andrews, 1♀ (CAFA). San Bernardino Co. : 4 mi E of Mentone, 750 m, May 11, 1978, R. T. Schuh and J. D. Pinto, Phacelia distans (Hydrophyllaceae) , 23 (AMNH). Bonanza Mine, Providence Mountains , E. P. Van Duzee, 13 (CAS). Kramer Hills, April 25, 1957, J. Powell, 13 (UCB). San Timoteo, May 6, 1948, R. A. Flock, Adenostoma sp. (Rosaceae) , 23 (UCR). San Diego Co. : 5 mi NE of Santa Ysabel , July 8, 1964, J. E. Prine, 2♀ (UCB). 8.2 mi E of Dulzura on Rt 94 at milepost 36.50, May 15, 1982, R. T. Schuh and B. M. Massie, Artemisia californica (Asteraceae) , 4♀ (AMNH). Borrego Springs, April 1, 1960, M. Wasbauer, Phacelia sp. (Hydrophyllaceae) , 43 (CAFA). Mt. Laguna , June 21, 1963, J. Powell, 13 (UCB). No specific locality, April 23, 1920, E. P. Van Du­ zee, 123, 14♀ (CAS). No specific locality, May 13, 1913, E. P. Van Duzee, 13 (CAFA). No specific locality, May 13, 1913, E. P. Van Duzee, 13 (CAFA). Pine Valley , June 13, 1927, C. C. Searl, 33 (SDNH). San Francisco Co. : No specific locality, paratypes: 33, 1♀ (CAS). San Francisco , May 13, 1927, C. L. Fox, 1♀ (USNM). San Francisco , Sand Dunes , May 23, 1947, C. L. Fox, 1♀ (CAS). San Luis Obispo Co. : 1 mi S of Cholame, April 30, 1963, H. B. Leech and P. H. Arnaud, Jr., 13 (CAS). 10 mi SE of Creston, April 25, 1968, J. Powell, 43, 3♀ (UCB). 18.5 mi E of Arroyo Grande , Huasna Valley , 1100 ft, May 9, 1985, R. T. Schuh and B. M. Massie, Salvia sp. (Lamiaceae) , 6♀ (AMNH). 3 mi W of Paso Robles, April 28, 1968, P. A. Opler, 43, 1♀ (UCB). 35 mi E of Santa Margarita , San Juan Creek , May 17, 1975, G. L. Parsons, 13 (OSU). 5 mi NE of Santa Margarita , May 16, 1980, J. D. Pinto, 1♀ (UCR). 5 mi S of Oceano, Oso Flaco Lake , May 11, 1965, J. Powell, 13, 1♀ (UCB). 6 air mi NE of Pozo , Black Mountain , 3300–3600 ft, May 1, 1962, C. A. Toschi, 13 (UCB). Arroyo Grande Creek SW of San Luis Obispo, 160 m, May 8, 1985, R. T. Schuh and B. M. Massie, 13, 4♀ (AMNH). La Panza Camp , April 25, 1968, Paul A. Opler, 53, 2♀ (UCB). near Atascadero , Morro Road , May 15, 1962, Brunson P. Bliven, 13, 2♀ (CAS). Ontario Hot Springs , April 22, 1922, E. P. Van Duzee, 183, 11♀ (CAS). Pozo , May 1, 1962, J. K. Drew, 13 (UCB). Shandon , May 14, 1961, Brunson P. Bliven, 13, 1♀ (CAS). Santa Barbara Co. : 1 mi S of Buellton, May 11, 1965, J. Powell, 23, 2♀ (UCB). Jalama Beach , April 23, 1966, J. Powell, 173, 6♀ (UCB). Los Prietos , April 23, 1966, R. L. Langston, 23 (UCB). Santa Cruz Island , Canyon Del Medio , May 1, 1969, D. S. Horning, 23, 5♀ (UCD). Upper Central Valley , Santa Cruz Island , April 26, 1966, J. Slater, Platystemon californicus (Papaveraceae) , 1♀ (UCB). Upper Oso Campground off Rt 154, 310 m, May 7, 1985, R. T. Schuh and B. M. Massie, Salvia leucophylla (Lamiaceae) , 63, 11♀ (AMNH). Shasta Co. : Castle Crag , June 25, 1941, Schuh and Grey , 1♀ (OSU). Hat Creek P. O., July 2, 1955, E. E. Lindquist, 2♀ (UCB). Hat Creek , July 4, 1955, D. L. Dahlsten, 1♀ (UCD). Sierra Co. : Downieville , July 8, 1952, M. Cazier, W. Gertsch, and R. Schrammel, 43, 8♀ (AMNH). Siskiyou Co. : 1 mi W of Bartle, 1220 m, July 7, 1979, R. T. and Joe Schuh , Monardella odoratissima (Lamiaceae) , 73, 5♀ (AMNH). 12.3 mi N of St. Hwy 89 on Powder Hill Road , July 19, 1985, G. M. Stonedahl and J. D. McIver, Chrysothamnus viscidiflorus (Asteraceae) , 13 (AMNH). Solano Co. : Vacaville , May 15, 1932, R. L. Usinger, 43 (UCB). Sonoma Co. : 2 mi E of Healdsburg, May 18, 1966, J. Powell, 2♀ (UCB). Hanleys , Mt. St. Helena , May 3, 1947, T. O. Thatcher, 3♀ (UCB). Stanislaus Co. : 2 mi W of Kyburz, June 11, 1966, W. Gagne, 2♀ (UCB). Del Puerto Canyon , Frank Raines Park , 1120 ft, May 15, 1971, C. B. Philip, 1♀ (CAS). Del Puerto Canyon , May 29, 1971, J. Marsh, 13 (UCD). Del Puerto Canyon, N Fork Del Puerto Creek , 1000 ft, June 12, 1975, R. Kawin, 2♀ (UCB). Tehama Co. : Red Bluff , Samson Slough , April 29, 1984, D. S. Chandler, 13 (UNHP). Trinity Co. : Carrville , 2400–2500 ft, June 4, 1934, G. and R. Bohart, 1♀ (CAS). Hayfork Ranger Station , 2300 ft, May 18, 1973, A. Chemsak, 1♀ (UCB). Snowslide Park , 7100 ft, July 30, 1972, T. Griswold, 1♀ (PUC). Tulare Co. : 4 mi NE of Lemon Cove, May 13, 1963, J. Powell, 43, 1♀ (UCB). 5 mi W of Three Rivers, Terminus Res. Camp , June 16, 1971, M. H. Sweet, 13, 1♀ (TAMU). Aukland , May 9, 1962, Brunson P. Bliven, 63, 2♀ (CAS). Badger , May 28, 1961, Brunson P. Bliven, 53, 9♀ (CAS). California Hot Springs , June 9, 1930, E. R. Lesch, 13, 1♀ (CAS). Isabella , May 6, 1931, E. C. Van Dyke, 1♀ (CAS). Sequoia National Park , May 28, 1949, W. Dwight Pierce, Calochortus luteus (Liliaceae) , 2♀ (LACM). Tuolumne Co. : 6 mi NE of Sonora, June 19, 1982, P. Oman, 2♀ (OSU). Sonora , May 21, 1969, A. E. and M. M. Michelbacher, 13, 1♀ (UCB). Ventura Co. : Blue Point Campground , vic. Piru Creek , May 7, 1966, Saul and Suzy Frommer , 23 (UCR). N end of Casitas Reservoir , March 15, 1967, P. A. Opler, 1♀ (UCB). Piru , April 20, 1932, E. P. Van Duzee, 133, 10♀ (CAS). Santa Paula, E. O. Essig , 13 (CAS). Santa Susana , May 27, 1930, 13, 1♀ (CAFA). Yolo Co. : 2 mi N of Rumsey, June 2, 1973, R. Harris, 1♀ (UCD). Davis , April 27, 1951, E. I. Schlinger, Arte­ misia vulgaris (Asteraceae) , 1♀ (UCD). Yuba Co. : Camp Far West Reservoir Dam , May 6, 1980, J. A. Powell, 13 (UCB). Dry Creek , Spenceville Wildlife Area , May 6, 1980, M. Buegler, 13, 1♀ (UCB). Idaho: Oneida Co. : Holbrook Summit , June 18, 1967, G. F. Knowlton, 13 (USU). Ireland Canyon , June 15, 1972, G. F. Knowlton, 13 (USU). Sayler Cow Camp , July 23, 1991, W. J. Hanson, 1♀ (USU). Nevada: Humboldt Co. : 14–16 mi N of Paradise Valley, 7100 ft, July 10, 1966, F., P., and M. Rindge, 13 (AMNH). Ormsby Co. : No specific locality, 73 (HELSINKI). No specific locality, July 1, 1900, C. F. Baker, 43, 2♀ (CAS). No specific locality, July 1, 1900, C. F. Baker, 53, 2♀ (HELSINKI). Oregon: Baker Co. : 9 mi E of Halfway, June 7, 1970, Oman , 13 (OSU). Coos Co. : 7.3 mi E of Bandon, 15 m, July 12, 1979, R. T. and Joe Schuh , 1♀ (AMNH). Grant Co. : 11 mi N of Seneca, May 14, 1973, Oman and Musgrave , 13 (OSU). Jackson Co. : 0.5 mi S of Siskiyou Summit on old Rt 99, 1350 m, June 26, 1979, R. T. and Joe Schuh , Artemisia cana (Asteraceae) , 93, 8♀ (AMNH). 0.5 mi S of Siskiyou Summit on old Rt 99, 1350 m, June 27, 1979, G. Stonedahl, Phacelia hastata (Hydrophyllaceae) , 33, 7♀ (AMNH). 0.5 mi S of Siskiyou Summit on Old Rt 99, 1350 m, June 27, 1979, J. D. Lattin, 1♀ (OSU). 0.5 mi S of Siskiyou Summit on Old Rt 99, 1350 m, June 27, 1979, M. D. Schwartz, Phacelia sp. (Hydrophyllaceae) , 213, 29♀ (AMNH). 12–15 mi E of Ashland, Dead Indian Road , 4500–4900 ft, July 17, 1930, H. A. Scullen, 13 (USNM). 12–15 mi E of Ashland, Dead Indian Road , 4500–4900 ft, June 7, 1970, H. A. Scullen, 13 (OSU). 29 mi ESE of Eagle Point, 4450 ft, June 23, 1978, Nancy L. Herman, 13 (AMNH). Fish Camp Post Office , in meadow, 1521 m, July 24, 1999, M. D. Schwartz, Lupinus sp. (Fabaceae) , 23, 7♀ (CNC). just E of Pinehurst, 1340 m, June 27, 1979, R. T. and Joe Schuh , Eriophyllum lanatum (Asteraceae) , 73, 10♀ (AMNH). Siskiyou Summit on I­5, July 4, 1982, G. M. Stonedahl and T. J. Henry, Phacelia hastata (Hydrophyllaceae) , 63, 2♀ (AMNH). Siskiyou Summit , old Siskiyou Hwy and Frontage Road , 4310 ft, July 1, 1994, M. D. Schwartz, Phacelia sp. (Hydrophyllaceae) , 43, 6♀ (CNC). Siskiyou , June 14, 1959, Kelton and Madge , 83, 7♀ (CNC). Klamath Co. : 13 mi W of Keno on Rt 66, 4600 ft, June 27, 1979, M. D. Schwartz, 13, 1♀ (AMNH). Chiloquin , Rt 97, July 4, 1982, G. M. Stonedahl and T. J. Henry, Plectritis sp. (Valerianaceae) , 53 (AMNH). Lake Co. : 5.7 of E of jct Rts. 395 and 140, Warner Mt. Pass, 1765 m, July 3, 1979, R. T. Schuh and B. Massie, 13, 1♀ (AMNH). Warner Canyon near Lakeview roadside, 2 mi E of Hwy 395, 5450 ft, July 19, 1971, J. D. Lattin, 93, 3♀ (OSU). Union Co. : La Grande, June 26, 1926, E. W. Davis, 1♀ (USNM) ; holotype male ( fusciflavus ) (USNM). Yamhill Co.: Dayton, Dorsey’s Gravel Bar , July 22, 1963, K. M. Fender, 23 (OSU). Wheatland Ferry , July 11, 1957, K. McKay­Fender, 13 (OSU). Utah: Emery Co. : Woodside , July 14, 1954, G. F. Knowlton, 13, 1♀ (KU). Washington: Grays Harbor Co. : Copalis Beach , July 6, 1978, Joe Schuh , 83, 3♀ (AMNH) .