Plagiognathus moerens Reuter, 1909: 80
publication ID |
https://doi.org/ 10.1206/0003-0090(2001)266<0001:RONWPF>2.0.CO;2 |
persistent identifier |
https://treatment.plazi.org/id/03F387FC-FF71-FF78-2D0A-FAD7FD2EFDB1 |
treatment provided by |
Felipe |
scientific name |
Plagiognathus moerens Reuter, 1909: 80 |
status |
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Plagiognathus moerens Reuter, 1909: 80 View in CoL (n. sp.). Chaetophylidea moerens: Knight, 1968: 33 (n.
comb.).
DIAGNOSIS: Recognized by the medium to large size, linear pattern of coloration with exocorium and clavus mostly opaque, pale, olivecolored, contrasting with dark endocorium (fig. 10), pronotum varying from entirely black to totally pale (olive), antennae entirely black (fig. 17), dorsal vestiture of recumbent, black, bristlelike setae, and surface of dorsum weakly polished (fig. 10). Linear pattern of coloration similar to lineatus (fig. 9) and shoshonea (fig. 13); easily distin guished from those species by the black, bristlelike setae on the dorsum and by the distinctive structure of the male genitalia (fig. 28), the vesica lacking a flange and with very long, evenly attenuated apical spines.
REDESCRIPTION: Male: Elongateovoid, moderate to large size; total length 3.84– 4.96, length apex clypeus–cuneal fracture 2.81–3.66, width across pronotum 1.23–1.46. COLORATION (fig. 10): Background coloration of dorsum deeply castaneous to blackish with longitudinal, opaque, pale (olive) areas; posterior margin of vertex pale at least at midpoint, sometimes across entire width, posterior lobe of pronotum often pale, costal margin of hemelytra dark or pale, exocorium almost totally pale, clavus pale except narrowly along scutellum and claval commissure, cuneus pale; corium narrowly pale adjacent to extreme base of membrane; membrane fumose, veins demarcating small cell and angle between posterior margin of small cell and posteromesial margin of cuneus pale; antennae entirely castaneous to black (fig. 17); labium castaneous; venter entirely castaneous, metathoracic scentgland evaporatory area sometimes partially pale; coxae dark, except occasionally pale at articulation with trochanters; trochanters dark; fore and middle femora weakly infuscate, hind femur heavily infuscate; tibiae weakly infuscate, hind tibiae sometimes more strongly so, spines of fore and middle tibiae without dark bases, spines of hind tibiae with small dark bases; tibiae dark at articulation with femora. SURFACE AND VESTITURE: Dorsum weakly granular, smooth, moderately shining. Vestiture of dorsum composed of nearly erect, black, bristlelike setae. STRUC TURE: Lateral corial margins weakly convex; frons distinctly tumid, clypeus visible from above; anteocular distance 2 times diameter of antennal segment 1; head projecting below eye by about 1.5 times diameter of antennal segment 1; labium reaching slightly beyond apex of hind coxae. GENI TALIA (fig. 28): Entire vesica very long, body of vesica relatively stout and strongly curving, more or less Jshaped, base of vesica falling slightly below level of secondary gonopore; apical spines long, of similar length, tapering gradually from gonopore to apex; no flange on vesica.
Female: Body shorter and more strongly ovoid than in male; pronotum and hemelytra more extensively pale than in male. Total length 3.84–4.96, length apex clypeus–cuneal fracture 2.81–3.66, width across pronotum 1.23–1.46.
HOSTS: Amsinckia spp. (Boraginaceae) , Phacelia spp. (Hydrophyllaceae) , and other herbaceous annuals.
DISTRIBUTION: From Idaho and Washington in the north, south through California and into Arizona.
DISCUSSION: Knight (1968) placed moerens Reuter in the new genus Chaetophylidea on the basis of its bristlelike vestiture. The vestiture is a generally variable attribute in Plagiognathus , and the situation in moerens certainly falls within the total range of variation. Because other attributes of moerens , including the male genitalia, fall well within the range of variation seen for the genus as conceived in this paper, I am treating Chaetophylidea as a junior synonym of Plagiognathus .
SPECIMENS EXAMINED: USA.— Arizona: Gila Co. : Rye, 3500 ft, April 18, 1982, D. A. and J. T. Polhemus, 1♀ (JTP). Maricopa Co. : Reavis Ranch Trail, 3600 ft, April 19, 1982, D. A. and J. T. Polhemus, 13 (JTP). Pinal Co.: Superior, April 16, 1928, A. A. Nichol, 13 (CNC). Yavapai Co.: 2 mi NW of Wickenburg on Rt 93, 625 m, April 1, 1981, R. T. Schuh and M. D. Schwartz, Amsinckia intermedia (Boraginaceae) , 603, 69♀ (AMNH). California: Alameda Co. : Corral Hollow, April 8, 1956, J. M. Burns, 13 (UCB). Amador Co.: Sutter Creek, April 24, 1985, 33, 6♀ (AMNH). Clear Lake Co.: Clear Lake, April 15, 1953, H. T. Osborn, Brassica sp. (Brassicaceae) , 2♀ (UCD). No specific locality, May 10, 1952, J. D. Lattin, 23 (AMNH). Fresno Co.: 12 mi W of Coalinga, April 8, 1951, E.G. Linsley, 13, 2♀ (UCB). 18 air mi SW of Mendota, Ciervo Hills, March 16, 1975, J. Powell and P.A. Rude, Amsinckia sp. (Boraginaceae) , 133, 8♀ (UCB). 5 mi S of Coalinga, Jacaitos Canyon, March 17, 1975, J. Powell, 23 (UCB). Fresno, April 10, 1963, Chester Latif, 1♀ (UCB). W of Herndon, San Joaquin River, April 3, 1970, J. Powell, Amsinckia (Boraginaceae) , 13 (UCB). Humboldt Co.: Dinsmores, June 7, 1942, B. P. Bliven, 63, 4♀ (CAS). Kern Co.: 1 mi E of Woody, May 3, 1964, P. Rude, 23, 5♀ (UCB). 1 mi NW of Tupman, March 19, 1975, J. A. Powell, Amsinckia sp. (Boraginaceae) , 23, 4♀ (UCB). 5 mi N of Lost Hills P.O., March 21, 1975, J. A. Powell, 13, 1♀ (UCB). 5 mi NE of Mojave, April 15, 1962, D. H. Janzen, Amsinckia sp. (Boraginaceae) , 63, 1♀ (UCB). 5 mi S of Democrat Springs, April 25, 1952, G. A. Marsh, 113, 12♀ (UCB). 7 mi SW of Maricopa , March 20, 1975, J. A. Powell, Amsinckia sp. (Boraginaceae) , 23 (UCB). Arvin, April 2, 1954, R. van den Bosch, 1♀ (UCR). Arvin, April 8, 1936, G. L. Smith, 13, 1♀ (UCB). Bakersfield, April 28, 1938, B. P. Bliven, 113, 14♀ (CAS). Cuyama Valley, April 8, 1932, E. P. Van Duzee, Amsinckia sp. (Boraginaceae) , 63, 6♀ (CAS). Edison, April 1, 1948, 2♀ (UCR). Havilah, 3000 ft, April 28, 1964, J. Powell, Plagiobothrys nothofulvus (Boraginaceae) , 23 (UCB). Maricopa , April 9, 1966, L. and C. W. O’Brien, 1♀ (UCB). Woody, NE of Bakersfield, April 16, 1961, B. P. Bliven, 63, 7♀ (CAS). Lake Co.: 5 mi NW of Middletown, June 4, 1963, W. Turner, 13 (UCB). Clear Lake Oaks, April 15, 1953, R. P. Allen, 13, 2♀ (CAFA). Clear Lake, May 10, 1952, J. D. Lattin, Raphanus sp. (Brassicaceae) , 33, 3♀ (OSU). Los Angeles Co.: 5 mi E of Llano, May 13, 1975, J. C. Hall, 1♀ (UCR). Calabasas, April 23, 1965, S. Seminoff, 13, 1♀ (CAFA). Claremont, Baker, 23, 1♀ (UCB, HELSINKI). Claremont, C. F. Baker, 293, 18♀ (CAS). Lancaster, June 8, 1962, M. E. Irwin, 13 (UCD). Mint Canyon, May 25, 1937, E. P. Van Duzee, 13 (CAS). No specific locality, March 22, 1939, K. E. Stag er, 13, 2♀ (LACM). San Clemente Island, E face Mt. Thirst, March 22, 1972, J. Powell, Amsinckia intermedia (Boraginaceae) , 33, 2♀ (UCB). Solemint, Mint Canyon, April 28, 1955, W. R. Richards, 123, 7♀ (CNC). Madera Co.: 4 mi NE of Madera, March 24, 1965, J. Powell, 13, 2♀ (UCB). Mendocino Co.: Hopland Field Sta., Kelsey Cab. Orchard area, 2500–2800 ft, May 14, 1977, T. and V. KeelerWolf, 43 (UCB). Hopland Field Station, April 28, 1977, R. L. Hanson, 13 (UCD). Merced Co.: 12 mi SE of Merced, April 13, 1964, R. P. Allen, 13 (CAFA). Modoc Co.: Waner Mts., Davis Creek, July 10, 1922, C. L. Fox, 13, 1♀ (CAS). Mon terey Co.: 3 air mi NE of Arroyo Seco guard station, Paloma Creek, 900 ft, May 4, 1975, 33, 3♀ (UCB). 5 mi NE of Arroyo Seco Guard Station, 800 ft, April 21, 1967, J.A. Powell, Amsinckia sp. (Boraginaceae) , 53, 1♀ (UCB). 6 mi W of Greenfield, Wiley Ranch, 1200 ft, May 2, 1975, S. L. Szerlip, J. A. Powell, and R. Wharton, Amsinckia sp. (Boraginaceae) , 173, 16♀ (UCB). Greenfield, March 12, 1967, P. Opler, 1♀ (UCB). Jolon, May 11, 1959, T. R. Haig, 23, 1♀ (UCD). San Ardo, April 29, 1953, R. P. Allen, Amsinckia sp. (Boraginaceae) , 43, 1♀ (CAFA). San Ardo, March 24, 1931, E. P. Van Duzee, 243, 8♀ (CAS). Summit of Carmel Divide, milepost 30.0, April 19, 1980, Russell and Schwartz, Hymenopappus filifolius (Asteraceae) , 273, 14♀ (AMNH). Orange Co.: Orange, 1♀ (LACM). Riverside Co.: 12 mi NW of Oak Grove on Hwy 79, April 5, 1969, S. Frommer et al., 93, 3♀ (UCR). 3 mi NW of Murietta, May 13, 1978, J. D. Pinto, Amsinckia sp. (Boraginaceae) , 4♀ (UCR). 3 mi W of Murriettta, Tenaja Road, 365 m, May 12, 1978, J. D. Pinto and R. T. Schuh, Amsinckia sp. (Boraginaceae) , 83, 36♀ (AMNH). 4 mi E of Elsinore, Red Rock Canyon, April 13, 1965, J. A. Powell, 73, 3♀ (UCB). 5 mi S of Sage, April 16, 1965, J. Powell, 23 (UCB). 6 mi SE of Corona, April 12, 1965, C. A. Toschi, 13 (UCB). 8 mi SE of Hemet, Bautista Canyon, May 1, 1974, J. D. Pinto, 13 (UCR). Big Springs Mts. near UCR campus, January 1, 1976, A. T. Mayor, 13 (UCR). Cactus Valley, T65 R1E Sec 8, April 29, 1980, J. N. Chandler, 13, 1♀ (UCR). Elsinore, March 3, 1974, C. Linn, 1♀ (UCR). Gavilan Hills, March 19, 1961, G. A. Gorelick, Amsinckia intermedia (Boraginaceae) , 13 (UCB). H. James Reserve, vicinity of Lake Fulmor, 5322 ft, May 22, 1976, B. A. Bowers, 1♀ (UCR). Lake Mathews, April 15, 1952, A. L. Melander, 13 (UCR). Lewis Valley, May 1, 1979, C. W. Melton, 23, 1♀ (UCR). Menifee Valley, hills on W end, 1800 ft, April 10, 1980 – April 21, 1983, J. D. Pinto, Cryptantha sp. (Boraginaceae) , 53, 1♀ (UCR). Millard Canyon, April 20, 1968, C. Beesley, Penstemon sp. (Scrophulariaceae) , 1♀ (UCR). Reche Canyon, May 12, 1963, E. I. Schlinger, 73, 1♀ (UCR). Riverside, March 18, 1965, M. E. Irwin, 13 (UCR). Riverside, March 28, 1935, H. L. McKenzie, Amsinckia douglasiana (Boraginaceae) , 73, 8♀ (UCD). Sage, April 15, 1965, J. Doyen, 43 (UCB). Temecula, April 23, 1951 – May 11, 1959, E. I. Schlinger, E. J. Taylor, 63, 1♀ (UCD). The Gavilan, May 17, 1951, E. I. Schlinger, 13 (UCD). Sacramento Co.: Citrus Heights, May 3, 1967, G. J. Keuter, 1♀ (CAS). Folsom, May 19, 1955, P.D. Hurd, Cryptantha sp. (Boraginaceae) , 13 (UCB). San Benito Co.: 10 mi S of Pinnacles Junction, California Hwy 25, April 16, 1966, C. W. O’Brien, 153, 18♀ (UCB). 5 mi S of Bitterwater, April 1, 1959, C. W. O’Brien, 13 (UCB). Big Panoche Creek, Fresno County line, April 21, 1967, P. A. Opler, Amsinckia (Boraginaceae) , 113, 11♀ (UCB). Mercy Hot Springs, April 6, 1951, E. J. Taylor, 9♀ (UCD). Pinnacles National Monument, April 19, 1980, M. D. Schwartz, 33, 4♀ (AMNH). Pinnacles National Monument, April 24, 1948, J. W. MacSwain, 13 (UCB). Pinnacles Post Office, April 6, 1951, E. J. Taylor, 33, 2♀ (UCD). San Bernardino Co.: 3 airline mi SE of New Idria, Clear Creek, 4000 ft, April 24, 1964, J. Powell, 23 (UCB). Apple Valley, May 12, 1955, W. R. M. Mason, 1♀ (CNC). Phelan, May 18, 1975, J. C. Hall, 8♀ (UCR). San Diego Co.: 2 mi NE of Lakeside, March 29, 1961, J. A. Powell, Lotus scoparius (Fabaceae) , 83, 14♀ (UCB). Mt. Laguna, July 5, 1963, H. L. Griffin, 1♀ (UCB). No specific locality, May 13, 1913, E. P. Van Duzee, 4♀ (CAS). San Luis Obispo Co.: 1 mi S of Cholame, April 30, 1963, P. H. Arnaud, Jr., 53, 5♀ (CAS). 10 mi SE of Creston, April 25, 1968, J. Doyen and J. Powell, 113, 6♀ (UCB). 10 mi W of Simmler, May 3, 1962, R. L. Langston, 33 (UCB). 12 mi E of Simmler, Tembler Range, 3200 ft, April 25, 1964, J. Powell, 2♀ (UCB). 12 mi NE of Pozo, La Panza Camp, April 29, 1962, J. K. Drew and P.D. Hurd, Cryptantha muricata (Boraginaceae) , 2♀ (UCB). 3 mi E of Pozo, May 1, 1962, P. D. Hurd, 23 (UCB). 3 mi N of Nacimiento Dam, April 14, 1967, P. Opler, 33, 2♀ (UCB). 3 mi N of Nacimiento, San Antonio Dam, April 14, 1967, P. A. Rude, 173, 13♀ (UCB). 5 mi NE of Santa Margarita, Jct. 58 & 229, April 12, 1967, P. A. Rude, 23 (UCB). 5.6 mi SW of Shandon, April 24, 1962, R.L. Langston, 1♀ (UCB). 6 mi SE of Creston, April 26, 1968, J. A. Chemsak, 1♀ (UCB). Atascadero, April 22, 1932, E. P. Van Duzee, 33, 3♀ (CAS). Cholame, April 11, 1960, E. G. Linsley, Phacelia sp. (Hydrophyllaceae) , 13 (UCB). La Panza Camp, April 26, 1968, J. Powell, 1♀ (UCB). Nacimiento Dam, April 14, 1967, P. A. Rude, 103, 7♀ (UCB). Pozo, April 28, 1962, C. A. Tosch, J. A. Powell, and J. K. Drew, 93, 20♀ (UCB). Simmler, March 20, 1940, J. W. Tilden, 13, 1♀ (CAS). SW of San Luis Obispo, Arroyo Grande Creek, 160 m, May 8, 1985, R. T. Schuh and B. M. Massie, 33 (AMNH). Templeton, March 22, 1931, E. P. Van Duzee, 23 (CAS). San Mateo Co.: Half Moon Bay State Beach, April 11, 1980, J. D. Pinto, 13 (UCR). Santa Barbara Co. : Cuyama, March 30, 1954, H. T. Osborn, Amsinckia sp. (Boraginaceae) , 23, 1♀ (CAFA). Santa Cruz Island, Central Valley, April 23, 1975, B. A. Bowers, 13 (UCR). Santa Clara Co. : Mt. Hamilton, April 15, 1947, G. Bohart, Gilia sp. (Polemoniaceae) , 13 (UCB). Pacheco Pass on Rt 152, 1380 ft, April 19, 1980, Russell and Schwartz, Hymenopappus filifolius (Asteraceae) , 33, 2♀ (AMNH). San Antonio Valley, April 20, 1948, Ray F. Smith, 13 (UCB). Shasta Co.: 7 mi S of Cottonwood, May 1, 1971, Oman, 23, 4♀ (OSU). Stanislaus Co.: Del Puerto Canyon, N Fork Del Puerto Creek, 1000 ft, April 13, 1980, N. Hostettler, 13 (UCB). Tehama Co. : 21 mi NW of Red Bluff on Rt 36, April 26, 1980, Russell and Schwartz, 13 (AMNH). Trinity Co.: Hayfork, 2300 ft, May 19, 1973, S. L. Szerlip, Phacelia heterophylla (Hydrophyllaceae) , 2♀ (UCB). Junction City, May 17, 1978, T. R. Haig, 13 (CAFA). Tulare Co.: 10.3 mi N of Lemon Cove, J21, March 25, 1981, M. D. Schwartz, Amsinckia sp. (Boraginaceae) , 123, 3♀ (AMNH). 11 mi NW of California Hot Springs , May 14, 1963, C. A. Toschi, 1♀ (UCB). 4 mi NE of Lemon Cove, May 13, 1963, J. Powell and S. W. Earnshaw, 2♀ (UCB). 6 mi N of Kaweah, April 28, 1979, J. Powell, 33, 1♀ (UCB). Lake Kaweak, Hwy 198, March 25, 1986, C. B. Barr, Amsinckia sp. (Boraginaceae) , 13, 3♀ (LSU). Lemon Cove, March 29, 1951, R.L. Usinger, Amsinckia sp. (Boraginaceae) , 13, 8♀ (UCB). Potwisha, 3 mi NE Ash Mountain Headquarters, 2000 ft, May 4, 1979, J. Powell, Amsinckia (Boragi naceae), 1♀ (UCB). Tuolumne Co.: 3 mi NE of Tuolumne, N Fork Tuolumne River, May 1, 1961, R. M. Brown, 5♀ (CAS). Ventura Co.: N end of Casitas Reservoir, March 15, 1967, P. A. Opler, 53, 5♀ (UCB). Piru, April 13, 1939, R. M. Bohart, 33 (UCB). Piru, April 20, 1932, E. P. Van Duzee, 1♀ (CAS). Quatal Canyon, NE Ventura Co., April 29, 1968, J. Powell, 43, 2♀ (UCB). Yolo Co.: Davis, June 6, 1942, E. I. Schlinger, 13 (UCD). Yuba Co.: 3 mi N of Smartville, Sierra Foothill Field Station, 1300 –1500 ft, May 2, 1980, W. Buegler, 13, 1♀ (UCB). Idaho: Clearwater Co.: Orofino, 1016 ft, June 17, 1935, 13 (CNC). Nez Perce Co.: Lewiston, 550 ft, May 11, 1935, W. E. Schull, 1♀ (CNC). Top of Lewiston Grade, June 30, 1975, J. Powell, Achillea sp. (Asteraceae) , 23 (UCB). Oregon: Gilliam Co.: 8 mi E of Olex, May 22, 1962, E. A. Dickason, 13 (OSU). Jackson Co.: 3 mi N of McLeod, McLeod State Park, Rogue River, 1550 ft, May 22, 1960, J. D. Lattin, 13, 1♀ (OSU). 3.2 mi S of Ruch, May 11, 1969, P. W. Oman, 1♀ (OSU). 8 mi N of Medford, TouVelle State Park, Rogue River, 1350 ft, May 21, 1960, J. D. Lattin, 43, 4♀ (OSU). north of Medord, Sam’s Valley, May 9, 1969, P. W. Oman, 13, 1♀ (OSU). TouVelle Park, May 2, 1970, Oman, 13 (OSU). Josephine Co.: 7 mi S of Grants Pass, Applegate River, May 18, 1962, 23, 3♀ (OSU). Sherman Co.: 5 mi W of Biggs Junction, Deschutes and Columbia Rivers , May 18, 1972, J. D. Lattin, 23, 2♀ (OSU). Umatilla Co.: Cayuse, April 23, 1939, J. Schuh and K. Gray, 13 (OSU). MiltonAthena, May 29, 1938, K. Gray and J. Schuh, 13 (OSU). Unknown Co.: Marlin, May 9, 1969, P. W. Oman, 1♀ (OSU). Wasco Co.: 5 mi E of Mosier, Mayer State Park, 550 ft, May 23, 1959, 1♀ (OSU). 5 mi N of Dufur, May 5, 1938, K. Gray and J. Schuh, 13 (OSU). 5 mi W of The Dalles, May 7, 1971, J. Sawbridge, 53, 2♀ (OSU). 9 mi E of The Dalles, May 13, 1972, Musgrave, 23 (OSU). The Dalles, April 30, 1938, K. Gray, 13 (OSU). Washington: Benton Co.: Hanford Site ALE, Snively Ranch, May 4, 1994, R. S. Zack, 13, 1♀ (CNC).
Plagiognathus monardellae , new species Figures 10 View Fig , 17 View Fig , 28 HOLOTYPE View Fig : Male : ‘‘[ USA] CALIFORNIA: Siskiyou Co. , 6.9 mi S of Medicine Lk. on
Powder Hill Rd., 19 July 1985, collectors GM Stonedahl and JD McIver, ex: Monardella odoratissima (Labiatae) ’’. Deposited in the American Museum of Natural History.
DIAGNOSIS: Recognized by the moderately large size, pale grayishgreen coloration (fig. 10), black antennae (17), relatively small eyes, ovoid body form, form of the male genitalia (fig. 28), and the preference for Monardella as the host. Similar in size and coloration of the dorsum to phaceliae (fig. 11), but that species with antennal segment 1 almost entirely pale and the remaining segments at most moderately infuscate (fig. 17). Also possibly confused with flavidus and guttatipes on the coloration of the dorsum, but those species with largely pale antennae.
DESCRIPTION: Male: Moderately large; total length 3.95–4.30, length apex clypeus– cuneal fracture 2.68–2.87, width across pronotum 1.11–1.15. COLORATION (fig. 10): General coloration, including most of venter and appendages, pale graygreen or yellowgreen; membrane weakly fumose, veins pale; antennae black (fig. 17) except for pale apical annulus on segment 1; apex of labium infuscate; thoracic sternum dark; abdominal venter variably infuscate; femora with some dark spots; dorsal tibial spines with small dark spots at bases; tibiae dark at femoral articulation. SURFACE AND VESTITURE: Dorsum weakly granular, smooth, weakly shining. Vestiture of dorsum composed of recumbent, pale, weakly golden, simple setae with some darker suberect setae on pronotum and anterolaterally on hemelytra. STRUC TURE: Body ovoid; frons moderately tumid as viewed from above, clypeus visible from above; anteocular distance 1.3 times diameter of antennal segment 1; head projecting below eye by 1.6 times diameter of antennal segment 1; labium reaching to near apex of hind coxae. GENITALIA (fig. 28): Vesica more or less Jshaped, body relatively slen der, base falling well below base of secondary gonopore; posterior apical spine elongate, slender, straight, very weakly angled relative to body of vesica, anterior spine smoothly curving and only slightly longer than posterior; flange developed, terminating at level of base of secondary gonopore.
Female: Shorter and more strongly ovoid than male; coloration similar to male. Total length 3.70–3.93, length apex clypeus–cuneal fracture 2.57–2.73, width across pronotum 1.11–1.20.
ETYMOLOGY: Named for its occurrence on Monardella spp.
HOST: Monardella odoratissima (Lamiaceae) .
DISTRIBUTION: Known only from the type locality in Siskiyou County, northern California.
PARATYPES: USA.— California: Siskiyou Co. : 6.9 mi S of Medicine Lake on Powder Hill Road, July 19, 1985, G. M. Stonedahl and J. D. McIver, 33, 12♀ (AMNH).
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