Plagiognathus alboradialis Knight, 1923: 439

SCHUH, RANDALL T., 2001, Revision Of New World Plagiognathus Fieber, With Comments On The Palearctic Fauna And The Description Of A New Genus (Heteroptera: Miridae: Phylinae), Bulletin of the American Museum of Natural History 2001 (266), pp. 1-267 : 35-36

Plagiognathus alboradialis Knight, 1923: 439 (n.

sp.).

DIAGNOSIS: Recognized by large size, elongate, slender body form, anterior pale portion of corium extending down radial vein, mostly pale costal vein and cuneus, and length of antennal segment 2 about 1.9 times width of head. General appearance very similar to the much more commonly collected obscurus ; distributions of these species broadly overlapping, in latter taxon anterior pale area of corium never extending down radial vein, and antennal segment 2 only about 1.7 times as long as width of head. Pattern of coloration in alboradialis (fig. 5) similar to flavoscutellatus , mundus , and paramundus in pale anterior portion of corium extending down radial vein; scutellum often pale laterally in alboradialis and apparently always so in paramundus . Former attribute also occurring in flavoscutellatus and latter known to occur in some populations of obscurus (fig. 10: obscurus 3). Distributions of alboradialis and paramundus not overlapping, clavus entirely dark in alboradialis but narrowly pale along claval suture in paramundus , and structure of genitalia distinctive for two species, with the posterior spine being slightly more slender and nearly as long as the anterior in alboradialis (fig. 20), and being slightly broader and reaching only to the level of the subapical bend of the anterior spine in paramundus (fig. 29); scutellum never pale medially as in flavoscutellatus (fig. 8).

REDESCRIPTION: Male: Elongate, more or less parallel­sided, large; total length 4.66– 4.99, length apex clypeus–cuneal fracture 3.25–3.38, width across pronotum 1.20– 1.31. COLORATION (fig. 5): Background coloration of dorsum castaneous, disc of pronotum sometimes weakly pale, scutellum sometimes pale laterally, corium pale on basal one­fourth, this area extending posteriorly along the radial vein to about midpoint of corium, costal vein almost entirely pale, cuneus usually entirely pale, corium narrowly pale adjacent to extreme base of membrane; membrane fumose, veins pale; all antennal segments castaneous to black except for pale, narrow, apical annulus on segment 1 (fig. 15); segment 1 and apex of labium castaneous, remainder pale; venter entirely castaneous, including metathoracic scent­gland evaporatory area; legs mostly pale yellowish, extreme base of coxae usually infuscate, femora with numerous dark spots; tibiae pale, dorsal spines with dark bases; tibiae dark at articulation with femora. SURFACE AND VESTITURE: Dorsum weakly granular, smooth, moderately shining. Face at and below base of clypeus more highly polished than remainder of body surface. Vestiture of dorsum composed of recumbent, golden, shining, simple setae. STRUCTURE: Lateral corial margins nearly straight; frons weakly convex, clypeus visible from above; anteocular distance nearly 2 times diameter of antennal segment 1; head projecting below eyes by 2 times diameter of antennal segment 1; labium reaching to about apex of hind coxae. GENITALIA (fig. 20): Body of vesica relatively stout and broadly curving, more or less U­shaped, base of vesica not quite reaching to level of secondary gonopore, posterior apical spine long, nearly straight, erect relative to body of vesica, anterior not as long as posterior, weakly angled relative to body of vesica, and angled near apex in lateral view; flange narrow and short, falling well below base of secondary gonopore and not extending beyond strap.

Female: Body shorter, broader, and much more strongly ovoid than in male (fig. 5). Vertex and most of pronotum pale, scutellum always pale laterally, leaving a dark, longitudinal, median stripe; hemelytra usually much more extensively pale than in male (fig. 5). Total length 4.31–4.70, length apex clypeus–cuneal fracture 2.98–3.26, width across pronotum 1.19–1.31.

HOSTS: Salix spp. (Salicaceae) ; also recorded from Rubus sp. (Rosaceae) .

DISTRIBUTION: New England and New York north to Nova Scotia, west to Colorado, Wyoming, and Idaho, and north to Alaska.

SPECIMENS EXAMINED: CANADA.— Alberta: Canmore , August 28, 1952, A. R. Brooks, 13 (CNC). Eisenhower Jct, August 5, 1970, L. A. Kelton, 23 (CNC). Elkwater Lake, July 20, 1956, O. Peck, 4♀ (CNC). Elkwater Park near campground stream, 3990 ft, July 16, 1990, M. D. Schwartz, Salix sp. (Salicaceae) , 13 (CNC). Elkwater Park, July 13, 1952 – July 21, 1952, L. A. Konotopetz and A. R. Brooks, 223, 11♀ (CNC). Elkwater Park, July 20, 1956, O. Peck, 73 (CNC). Jasper, August 29, 1970, L. A. Kelton, 13, 1♀ (CNC). Kananaskis Hiway, July 25, 1975, L. A. Kelton, Salix sp. (Salicaceae) , 53, 3♀ (CNC). Lake Louise, August 3, 1970, L. A. Kelton, 1♀ (CNC). McMurray, August 16, 1953, G. E. Ball, 23, 3♀ (CNC). Nordegg, July 3, 1921, J. Mc­ Dunnough, 1♀ (CNC). Slave Lake, August 14, 1924, O. Bryant, 13 (USNM). Waterton, July 26, 1972, L. A. Kelton, 13, 1♀ (CNC). Labrador: Cartwright, August 2, 1955, F. E. Sterns, 13, 1♀ (CNC). Manitoba: 10 mi W of Roblin on Rt 5, July 15, 1954, Brooks and Wallis, 23, 1♀ (CNC). Boissevain, July 20, 1953, Brooks and Kelton, 13 (CNC). Brandon, July 18, 1958, R. L. Hurley, 23 (CNC). Churchill, August 6, 1952, J. R. Chillcott, 1♀ (CNC). Falcon Lake, June 25, 1972, L. A. Kelton, 23 (CNC). Horton, July 25, 1953, Brooks and Kelton, 13, 4♀ (CNC). Oak Lake, July 1, 1953, Brooks and Kelton, 1♀ (CNC). Russell, July 21, 1954, Brooks and Wallis, 13 (CNC). Turtle Mt. Forest Reserve, Internation Peace Gardens, July 17, 1958, J. G. Chillcott, 13 (CNC). Turtle Mt., July 22, 1953, Brooks and Kelton, 123, 18♀ (CNC). Virden, July 8, 1953, Brooks and Kelton, 13 (CNC). New Brunswick: Bathurst, June 28, 1966, L. A. Kelton, Rubus sp. (Rosaceae) , 73, 3♀ (CNC). Cambellton, June 29, 1966, L. A. Kelton, Rubus sp. (Rosaceae) , 83, 7♀ (CNC). Edmunston, July 1, 1966, L. A., Kelton, Salix sp. (Salicaceae) , 23 (CNC). Fundy National Park, July 6, 1966, L. A. Kelton, 13 (CNC). Petersville, July 5, 1966, L. A. Kelton, 13 (CNC). Woodstock, June 22, 1966, L. A. Kelton, 1♀ (CNC). Newfoundland: Spruce Brook, August 8, 1912, 23, 9♀ (AMNH). Northwest Territories: Aklvik, July 16, 1931, Bryant, 13 (CAS). Nova Scotia: Kentville, July 3, 1966, L. A. Kelton, 13 (CNC). Kentville, July 7, 1924, R. P. Gorham, 13 (CNC). Mt. Uniacke, July 12, 1966, L. A. Kelton, 33 (CNC). Truro, July 12, 1913, R. Matheson, 13 (USNM). Ontario: Collingwood, June 13, 1962, Kelton and Thorpe, 13 (CNC). Dunnville, July 9, 1962, Kelton and Thorpe, 3♀ (CNC). Mazinaw Lake, June 20, 1974, D. G. Reid, 13 (CNC). Norway Point, Lake of Bays, June 28, 1922, J. McDunnough, 13 (CNC). Parry Sound, July 10, 1915, paratype: 13 (USNM). Thessalon, July 11, 1965, W. Gagne, 23 (UCB). Quebec: Duchesnay, July 5, 1953, R. Lambert, 13 (CNC). Fabre, July 12, 1963, W. Gagne, 1♀ (CNC). Frelighsburg, June 21, 1972, L. A. Kelton, 13 (CNC). Great Whale River, August 29, 1949, J. R. Vockeroth, 1♀ (CNC). Knowlton, July 11, 1929, L. J. Milne, 13 (CNC). Lac Mondor, Ste. Flore, July 6, 1951, E. G. Munroe, 23 (CNC). Laniel, July 1, 1963 – July 16, 1963, W. Gagne, 373, 18♀ (CNC). Mt. Albert, July 11, 1954, W. J. Brown, 13 (CNC). Quinze Lake, August 18, 1907, W. J. Palmer, 13 (CAS). Yarms, July 23, 1955, C. H. Mann, 1♀ (CNC). Kamouraska Co.: Parke Reserve, July 11, 1957, G. E. Shewell, 23, 2♀ (CNC). Missisquoi Co.: Mt. Pinacle, 8 km E of Frelighsburg, 400 m, June 20, 1991, M. D. Schwartz, 13 (CNC). Saskatchewan: Elbow, July 1, 1960, A. R. Brooks, 13 (CNC). Kenosee, July 19, 1958, A. and J. Brooks, 103, 12♀ (CNC). Lake Madge, July 12, 1954, Brooks and Wallis, 23, 2♀ (CNC). Prince Albert, July 23, 1959, A. and J. Brooks, 33 (CNC). Saskatoon, June 26, 1951, L. A. Konotopetz, 23, 2♀ (CNC). White Forest, July 24, 1950, L. A. Konoto­ petz, 13 (CNC). USA.— Alaska: Fort Yukon, July 15, 1951, R. I. Sailer, 23 (USNM). Colorado: Jackson Co.: 1 mi E of Gould, August 11, 1969, J. C. Schaffner, 13 (TAMU). Larimer Co.: 46 mi W of Fort Collins, Bennett Crk. Pic. Grd., Pingree Pk. Rd., July 14, 1986, R. T. Schuh and J. T. Polhemus, 1♀ (AMNH). Idaho: Blaine Co.: Bellveue, July 30, 1932, W. E. Shull, 23 (USNM). Maine: Cumberland Co.: Portland, July 9, 1909, E. P. Van Duzee, 1♀ (CAS). Oxford Co.: near Bethel, July 7, 1900, B. Banks, 33, 4♀ (AMNH). Piscataquis Co.: Traveller Mountain, July 27, 1910, paratype: 13 (CAS). Capens, July 20, 1907, paratype: 13 (CAS). Washington Co.: Calais, July 10, 1900, 1♀ (CAS). Eastport, July 15, 1909, paratypes: 13, 2♀ (CAS). Machias, July 22, 1909, paratype: 43, 2♀ (CAS). Princeton, July 12, 1907, paratypes: 13, 1♀ (CAS). Massachusetts: Norfolk Co.: Wellesley, July 11, 1909, E. P. Van Duzee, 13 (CAS). Minnesota: Itasca Co.: Deer Lake, June 15, 1986, D. A. Rider, 33, 1♀ (DAR). Montana: Gallatin Co.: Bozeman, July 21, 1985, R. Styles, 13 (DAR). New Hampshire: Carroll Co.: Crawford Notch, July 23, 1900, N. Banks, 13 (AMNH). Coos Co.: Bretton Woods, June 30, 1909, E. P. Van Duzee, 13 (CAS). Glen House, July 15, 1915, paratypes: 63, 2♀ (CAS). Mount Washington, Halfway House, July 18, 1915, paratype: 13 (CAS). Mount Washington, July 4, 1915, 2♀ (CAS). Pinkham Notch, July 6, 1900, N. Banks, 13 (AMNH). Grafton Co.: Hubbard Brook Exp. Forest, July 10, 1975, M. Schwartz, 1♀ (AMNH). New York: Erie Co.: Elma, June 25, 1911, E. P. Van Duzee, 1♀ (CAS). Hamburg, July 23, 1905, E. P. Van Duzee, 13 (CAS). Essex Co.: Lake Placid, July 19, 1962, J. R. Vockeroth, 13 (CNC). Lake Placid, September 22, 1902, E. P. Van Duzee, 13 (CAS). Whiteface Mountain, August 22, 1916, H. H. Knight, 1♀ (CAS). Genesee Co.: Batavia, June 25, 1915, H. H. Knight, paratypes: 23 (CAS). Hamilton Co.: Speculator, July 20, 1909, E. P. Van Duzee, 13 (CAS). St. Lawrence Co.: Cranberry Lake, July 11, 1917, C. J. Drake, 13, 1♀ (TAMU). North Dakota: Ransom Co.: 4 mi NW of McLeod, June 25, 1992, D. A. Rider, 13 (DAR). Traill Co.: No specific locality, August 9, 1923, A. A. Nichol, 2♀ (USNM). Vermont: Washington Co.: Montpelier, June 23, 1900, C. W. Johnson , 13 (AMNH). Montpelier, June 25, 1906, paratypes: 13, 1♀ (CAS). Wyoming: Park Co.: Yellowstone National Park, July 20, 1920, A. A. Nichol, 13 (USNM).