Tortopsis unguiculatus (Ulmer) Ulmer, 2010

Molineri, Carlos, 2010, A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae), Zootaxa 2481, pp. 1-36 : 23

Tortopsis unguiculatus (Ulmer) New combination

Campsurus unguiculatus Ulmer 1920: 4 .

Tortopus unguiculatus ; Ulmer 1942: 109; Traver 1950: 600.

Material ( ZMH): 3 male labeled “ Type ”, “Columbien/Behn vend. 3.XII.1900.”, “G. Ulmer determ./ 1916– 1923 ”.

Male imago. General coloration faint in the old pinned types. Genitalia: long and cylindrical penes completely separated from their bases ( Fig. 41 ); sclerotized spine on inner apical margin with wide rounded base and slightly hooked apex ( Fig. 43 ); parastyli long and acute, markedly curved dorsally, the main axis of the basal third, form a 90° angle with the axis of the distal third ( Fig. 42 ); parastyli conical and entire, not furrowed longitudinally ( Fig. 41 ).

Discussion and diagnosis. Ulmer (1920) described this species from 3 male imagos from Colombia, and later ( Ulmer, 1942) provided a description of a female imago from Colombia and cited some male imagos from Reventazón ( Costa Rica). Additional records exist for this species in Mexico, Nicaragua (Zelaya) and Costa Rica ( Maés 1988) and Guatemala and Honduras ( Lugo-Ortiz & McCafferty 1996), but this material should be revised with the new data offered here to confirm its specific identity. Tortopsis unguiculatus can be distinguished from all other species of the genus by: 1) fore wing length 9.0–11.0 mm (male), 11.0–12.0 mm (female); 2) wings hyaline translucent, veins whitish translucent; 3) parastyli long and very strongly curved ( Fig. 42 ); 4) penes cylindrical, apical spine with wide base and a small hook ( Fig. 43 ).

Figures

FIGURES 41 – 43. Tortopsis unguiculatus, type male genitalia, photographs and interpretations (line drawings). 41, ventral view; 42, lateral view; 43, laterodorsal view (details of apex of penes).