Lithocorydalus , Jepson, James E. & Heads, Sam W., 2016

Jepson, James E. & Heads, Sam W., 2016, Fossil Megaloptera (Insecta: Neuropterida) from the Lower Cretaceous Crato Formation of Brazil, Zootaxa 4098 (1), pp. 134-144: 139

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gen. nov.

Genus: Lithocorydalus  gen. nov.

Etymology. From lithos, Greek for stone and Corydalus  an extant genus of megalopteran.

Type species: Lithocorydalus fuscata  gen. et sp. nov.

Diagnosis. Lithocorydalus  gen. nov. differs from the other genera of Corydalidae  by a combination of the following characters: Elongate forewing: length 17.9 mm. Very dark wing colouration. Venation relatively simple: Rs branches majority simple, MA simple, MP deeply forked: MP 1, MP 2 simple. Elongate hind wing, free branch of MA in hind wing, forming long cell between it and R. Two nygmata present, tibial spur absent.

Comparison. The absence of a bilobed 4 th tarsomere in Lithocorydalus  gen. nov. separates it from taxa in Sialidae  and Corydasialidae  . Lithocorydalus  gen. nov. can be separated from Parasialidae  and further separated from Sialidae  by its general wing venation. Lithocorydalus  gen. nov. has a relatively small to medium sized forewing, which is larger than Cretochaulus (10 mm), but smaller than the other taxa: Cratocorydalopsis  gen. nov (25 mm), Eochauliodes (30 mm), and Jurachauliodes (35 mm). The uniform, very dark colour pattern differs from all other genera: Cratocorydalopsis  gen. nov.: (dark base and apex, pale centre), Eochauliodes (hyaline with dark stripes), Jurachauliodes (hyaline with some dark patches). The venation is relatively simple in Lithocorydalus  gen. nov. compared to the other genera, with regards to Rs, Cratocorydalopsis  gen. nov differs by having more branches of Rs; in Cretochaulus and Eochauliodes, Rs is dichotomously forked, and in Jurachauliodes Rs has three branches that are all forked in forewing, but in the hind wing, all but one is forked. The structure of MA is similar to Cretochaulus, which is also simple; in all other genera it is forked. The deeply forked MP is shared with all genera, but the simple MP 1 and MP 2 differs from Cretochaulus, Eochauliodes and Cratocorydalopsis  gen. nov., which have MP 1 forked and MP 2 simple, Jurachauliodes has MP 1 forked twice (forewing), dichotomously forked (hind wing), MP 2 simple (forewing), and forked (hind wing). The elongate hind wing is observed in Eochauliodes, but broad in all other genera. The free branch of MA is similar to the other genera, except Eochauliodes, which is absent (possibly due to preservation). The presence of two nygmata is the same in the other fossil genera except Jurachauliodes and Cratocorydalopsis  gen. nov.: in which it is absent, however this may be due to preservation. The absence of a tibial spur is also observed in Cratocorydalopsis  gen. nov.; in Cretochaulus it is unknown, but in Eochauliodes and Jurachauliodes it is present.