Satsuma, Adams, 1868

Hwang, Chung-Chi & Wu, Shu-Ping, 2024, Three new species of Satsuma Adams, 1868 (Gastropoda, Camaenidae) from Taiwan, Zoosystema 46 (3), pp. 61-76 : 73-74

KEY TO THE DEXTRAL SPECIES OF SATSUMA ADAMS, 1868 IN TAIWAN

1 Shell high conic; shell height> shell width .................................................................................................. 2

— Shell depressed conic; shell height <shell width .......................................................................................... 3

2 Shell white or yellowish golden; with/without polymorphic bands ................................................................. ................................................................................................ S. albida species complex (see Wu et al. 2008)

— Shell brown; with a peripheral band ............................................................... S. sphaeroconus (Pfeiffer, 1866)

3 Shell colour yellow .................................................................................................... S. bacca (Pfeiffer, 1866)

— Shell colour light brown to dark brown ....................................................................................................... 4

4 Umbilicus totally covered by reflexed columellar lip .................................................................................... 5

Umbilicus partly open to completely open .................................................................................................. 6

5 Shell large-sized, shell width> 50 mm ................................................................... S. arisana ( Kuroda, 1941)

Shell medium-sized, shell width <45 mm ................................................................... S. tani ( Kuroda, 1941)

6 Shell width> 35 mm ................................................................................................................................... 7

Shell width <35 mm ................................................................................................................................. 10

7 Upper lip clearly sinuous; fold on basal lip present ............................................... S. bairdi (H. Adams, 1866)

Upper lip smoothly curved; fold on basal lip absent .................................................................................... 8

8 Spire lower; shell height/shell width <0.7 ............................................................ S. takahasii ( Kuroda, 1941)

— Spire higher; shell height/shell width> 0.7 .................................................................................................. 9

9 Umbilicus completely open; penial caecum weak ........................ S. longkiauwensis Wu, Lin & Hwang, 2007

— Umbilicus partly open; penial caecum low conical .................................................................. S. nivnu n. sp.

10 Periphery angulate to carinate ................................................................................................................... 11

— Periphery round, at most roundly angulate ................................................................................................ 16

11 Umbilicus not covered by reflexed columellar lip ............................................. Satsuma mellea (Pfeiffer, 1866)

— Umbilicus partly covered by reflexed columellar lip ................................................................................... 12

12 Periphery sharply carinate; surface immediately below periphery concave; basal lip flat .................................. ............................................. Satsuma friesiana (Möllendorff, 1884) + syn. Coniglobus carinatus Chang, 1990

— Periphery bluntly carinate; surface immediately below periphery convex; basal lip smoothly curved .......... 13

13 Periphery angulate in front; whorls flat to weakly convex .......................................................................... 14

— Periphery round in front; whorls convex ................................................................................................... 15

14 Whorls flat; base convex; axial striae dense ........................................ S. succincta succincta (H. Adams, 1866)

— Whorls convex; base less convex; axial striae loose ............................................... S. paiwanis ( Kuroda, 1941)

15 Spire higher; shell height/width ratio = 0.79; peristome very thick; whorls less convex ................................... ........................................................................................................... S. succincta rubrotincta ( Kuroda, 1941)

— Spire lower; shell height/width ratio = 0.69; peristome not particularly thickened; whorls convex .................. ............................................................................................................ S. succincta amblytropis (Pilsbry, 1901)

16 Umbilicus close ................................................................................................ S. takkiriensis ( Kuroda, 1941)

— Umbilicus partly covered ............................................................................................................................ 17

17 Basal lip curved ............................................................................................... S. litus (Chang & Tada, 2000)

— Basal lip flat ................................................................................................................................................ 18

18 Upper surface roughly and loosely ridged; outer lip sinuate ........................................................................ 19

— Upper surface finely and densely ridged; outer lip smoothly curved ............................................................ 20

19 Basal lip horizontal, straight; surface above periphery malleated ........................... S. nux ( Möllendorff, 1888)

— Basal lip straightly downwrds; surface above periphery not malleated ................................ S. sunlinksea n. sp.

20 Spire high, shell height/shell width> 0.8 ................................................................................................... 21

— Spire low, shell height/shell width <0.75 ................................................................................................... 22

21 Upper lip smoothly curved; basal lip horizontal with a weak fold; penial pouch absent .................................. .............................................................................................................................. S. sericata ( Kuroda, 1941)

— Upper lip sinuous; basal lip oblique without a fold; penial pouch present .......................... S. hsuehshan n. sp.

22 Tooth on basal lip apparent; flagellum hooked on tip, tapering ............................ S. chengi Wu & Tsai, 2015

— Tooth on basal lip inapparent; flagellum spirally tapering ........................................................................... 23

23 Distinguished by molecular characters only; shell larger; shell colour paler; penis shorter; distributed around Chipen, southeastern Taiwan.................................................................................... S. mii Wu & Wu, 2017

— Distinguished by molecular characters only; shell smaller; shell colour darker; penis longer; distributed around Antong, eastern Taiwan.................................................................................... S. paradoxa Wu & Tsai, 2016