Glyphiulus subgranulatus, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group, Zoosystema 29 (1), pp. 7-49 : 21-23

publication ID	https://doi.org/ 10.5281/zenodo.5392079
DOI	https://doi.org/10.5281/zenodo.5468294
persistent identifier	https://treatment.plazi.org/id/03F4C378-A221-3058-FF45-0270FB77F935
treatment provided by	Marcus
scientific name	Glyphiulus subgranulatus
status	sp. nov.

Treatment

Glyphiulus subgranulatus View in CoL n. sp.

( Figs 7-9 View FIG View FIG View FIG )

TYPE MATERIAL. — China. Yunnan Prov., Mengzi County, cave near footpath to plateau, 5.I.1989, leg. P. Beron, holotype ♂ ( NMNHS) ; paratypes 7 ♂♂, 14 ♀♀, 11 juv. ( NMNHS) ; 2 ♂♂, 2 ♀♀ ( MNHN GA 035 View Materials ) ; 2 ♂♂, 2 ♀♀ ( ZMUM) ; 1 ♂, 1 ♀ ( ZMUC) ; 1 ♂, 1 ♀ ( MCSNV) ; 1 ♂, 1 ♀ ( SEM). — Yunnan Prov., Mengzi County, Pothole No. 2 ( Ma Fa Tiao Dong ), 6.I.1989, leg. P. Beron, paratype 1 ♀ ( NMNHS) .

ETYMOLOGY. — To emphasize the obvious similarity to G. granulatus .

DIAGNOSIS. — Differs from the most similar congeners by the crests on the collum being underdeveloped (especially so anteriorly), the body segments somewhat compressed laterally, the anterior gonopods with unusually slender telopodites, etc.

DESCRIPTION

Length of adults of both sexes 19-28 mm, width 1.0- 1.3 mm; males usually a little shorter and more slender than females. Coloration much as in G. granulatus , variegate, yellow-brown, with a dark brown vertex, blackish ocellaria, mainly brown crests on collum and grey-brownish lateral longitudinal stripes running down from dark brown ozoporiferous tubercles; dorsum generally light, but a thin axial line mostly present due to darker median crests.

Adults with 38-52p+5-2a+T in males, 39-63p+5- 1a+T in females; largest juvenile ♂ with 31p+3a+T; two smallest juveniles with 17 or 18 haplopodous segments after 6th, plus 2a+T. Holotype about 23 mm long, 1.0 mm wide, with 47p+2a+T.

Other characters as in G. granulatus , including mentum either divided (n = 1) ( Fig. 9B View FIG ) or undivided (n = 1) ( Fig. 7A View FIG ), 11-18 ocelli, antennae ( Figs 8A View FIG ; 9A View FIG ), the carinotaxy formulae ( Fig. 7 View FIG B-F), claw, posterior gonopod structure ( Fig. 9F, G View FIG ), male legs 1 and 2 ( Figs 8D View FIG ; 9C, D View FIG ), telson ( Fig. 8B View FIG ), etc., but body size much larger (see above); segments compressed laterally ( Fig. 8C View FIG ); crests on collum (7+m+7 at caudal margin) evidently reduced in size, almost obsolete anteromedially ( Figs 7B View FIG ; 8A View FIG ); legs somewhat longer (about 4/5 as long as body height); anterior gonopods with telopodites more slender, while the coxosternum has much shorter distomedian outgrowths and acute distolateral corners ( Fig. 9E View FIG ).