Manurevana , Hoch, Hannelore, 2006

Hoch, Hannelore, 2006, New Cixiidae from Eastern Polynesia: Oteana gen. nov. and Manurevana gen. nov. (Hemiptera: Fulgoromorpha), Zootaxa 1209, pp. 1-47: 41-42

publication ID 10.5281/zenodo.172430

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gen. nov.

Manurevana  gen. nov.

Type species: Manurevana draconarius ( Fennah, 1958)  comb.nov. Oliarus draconarius Fennah, 1958: 130 


Small inconspicuously colored cixiids with depressed body, mesonotum pentecarinate, tegmina shallowly tectiform.


Head (fig. 91)

Vertex ca. 1.4 times as long as posteriorly wide, in dorsal aspect surpassing the anterior margin of the compound eyes with ca. 1 / 3 its total length; lateral carinae more or less parallel in posterior half, anteriorly converging, ridged; median carina present in posterior half of vertex; posterior margin of vertex deeply incised. Areolets medially divided by a short, obtuse carina. Transition frons/vertex rounded, without a distinct transverse carina. Frons longish, narrow, ca. 0.8 times as wide as medially long, widest at level of the antennae. Lateral ocelli present, distinct; median frontal ocellus absent. Frontoclypeal suture more or less straight. Frons and clypeus with a ridged median carina, frontal carina forked towards apex. Rostrum attaining metatrochanters. Antennae small, not visible in dorsal aspect, 1 st antennal segment short, ring­like, 2 nd antennal segment subglobose, irregularly beset with sense organs (plaques).


Pronotum short, posterior margin deeply incised, ca. 1.2 times wider than head, with distinct median and lateral carinae. Mesonotum pentecarinate, about as wide as medially long. Tegmina and wings well developed. Tegmina hyaline, translucent, shallowly tectiform, in repose surpassing tip of abdomen with ca. 1 / 3 their total length. Tegmen ca. 2.5 times longer than maximally wide, apex of outer subapical cell slightly proximad of apex of inner subapical cell, intercubital transverse veinlet entering margin of tegmen distinctly distad of claval suture, longitudinal veins inconspicuously granulate. Metatibiae laterally with 3 strong spines, distally with (5 –) 6 spines, arranged in an oblique row, lateral spine more strongly pronounced. Metabasitarsus only slightly longer than 2 nd and 3 rd metatarsal joints together, distally with (6 –) 7 spines; 2 nd metatarsal joint distally with (7 –) 8 spines. Metatarsal joints without macrochaetae (platellae).

Male genital complex.

Genital segment bilaterally symmetrical; posterior margin in lateral aspect broadly rounded; medioventral process helmet­shaped, produced with a sharp ridge posteriorly into a pointed tip, dorsally rounded. Anal segment hood­shaped, bilaterally symmetrical, longish ovate, ventrally concave. Parameres in lateral aspect wide at base, distally narrowing and apically broadening into a bluntly triangular portion, medially bearing a strongly developed ridge. Aedeagus. Shaft with two spinose processes. Flagellum long, in repose reaching base of shaft.

Female genitalia.

Caudal margin of 7 th sternite shallowly sinuate, medially slightly convex, produced into a shallow lobe. Ovipositor short, stout, in repose directed straight caudad. 9 th tergite truncate, caudally concave, with wax producing area distinct, medially not divided. Anal segment longish ovate, with lateral margins nearly parallel, narrow at base, ventrally not concave.


Named for the type locality, Mt. Manureva on Rurutu. The gender is masculine.


Manurevana  gen.nov. can be distinguished from other pentastirine genera by bodily proportions, shape and arrangement of spines of the metatarsi (7, 8, no platellae), and the configuration of the male genitalia.