Ceratojoppini HEINRICH

A. M, 2009, Illustrated key to the tribes of subfamilia Ichneumoninae and genera of the tribe Platylabini of world fauna (Hymenoptera, Ichneumonidae), Linzer biologische Beiträge 41 (2), pp. 1317-1608 : 1343-1345

Tribe Ceratojoppini HEINRICH

Ceratojoppini HEINRICH 1938 - Mem. de l'Acad. Malgache 25: 121.

Type genus: Ceratojoppa CAMERON.

Ceratojoppini : HEINRICH 1968 - Ichn. Stenop. of Africa 2: 201.

D i s t r i b u t i o n: South of Africa and Madagascar.

I n t r o d u c t i o n:

Tribe Ceratojoppini is one of the most distinctly detached tribe of the subfamily. The peculiarities of morphology of different parts of the body have characters which characteristic

to others subfamilies of Ichneumonidae (for example, morphology of propodeum and first tergites of abdomen are similar to Ophioninae ).

Very long and slender flagellum, head with very broad genae and slightly differentiated (but distinct) fields of face and clypeus, short and broad mandibles, with teeth divided by deep broad gape and frons, with longitudinal carinae are character to the tribe. The lateral lobes of mesonotum along the full length are separated from elevated middle lobe by irregular and roughly transversally-wrinkled notauli. Propodeum is long, convex from lateral, without distinct fields, its dorsal surface is only with single transverse carina, separating smooth front part from roughly-wrinkled hind one. Legs are slender, long. Wings are with quadrangular, sometimes petiolate areolet. Abdomen of females from above is slen- der, long, with long second tergite, oxypygous. Thyridia of second tergite are very large, moved from base of tergite, with narrow interval between them.

M o r p h o l o g y:

F l a g e l l u m: Very long and slender; of females, bristle-shaped, ventrally flattened beyond middle and sometimes slightly widened, with very strongly elongated basal segments, sharply attenuated toward apex. Flagellum of males, not nodose with the exception of apex, with row extremely narrow, slightly expressed bacilliform tyloides. Flagellum almost equal by length to front wing or longer.

H e a d: Head contour from front roundly narrowed downwards; genae from lateral broad, temples from above roundly narrowed behind eyes (head from above not stout). Occipital carina high; mandibles short, broad, sharply curved and carinated with large teeth separated by wide gape between teeth which situated in the same plane; clypeus, with oblique lateral sides, rounded corners and straight front margin; labrum slightly project from under clypeus; clypeal foveae large, deep; fields of face slightly differentiated, but distinct, middle field only just convex; antennal cavities deeply impressed, of moderate size; frons with five longitudinal carinae of specific morphology; ocellar triangle elevated and surrounded by deep groove; sculpture of head mainly coriaceous or polished.

T h o r a x: Pronotum with distinct epomiae; lower angle of pronotum with very sharp angle (African species). Mesonotum slightly convex, middle lobe elevated, with flattened front surface, lateral lobes of mesonotum strongly lowered, theirs external margins form elevated ribs directed mainly to apex; notauli sharp all round, forming all round broad shallow furrows, roughly and irregularly transversally-wrinkled; prepectus above front coxae with two longitudinal ribs (at least of Ceratojoppa cornuta CAMERON ); subalarum thin, sharp; area of mesopleural fovea only just impressed; scutellum high elevated above postscutellum, with high lateral and sometimes apical carinae, dorsally flat or convex. Hind margin of metanotum with triangular projections. Propodeum long, from lateral gradually slanting to apex; basal furrow distinct; area basalis and areae superoexternae merged to one, usually smoothed transversal basal area, limited basally and apically by distinct carinae; carina metapleuralis distinct, other carinae atrophied; spiracles long; propodeum from apical margin of smoothed basal area roughly reticulate-wrinkled; apices of areae dentiparae sometimes slightly indicated.

L e g s: Quite long, thin; femora extremely thin, coxae comparatively small; hind coxae of females sometimes with distinct scopa.

W i n g s: Areolet quadrangular, to petiolate; nervulus interstitial or slightly postfurcal;

radius basally sinuous; membrane of wing hyaline, sometimes wings apically darkened. Wings distinctly shorter than body length.

A b d o m e n Abdomen of females from above slender, long; second tergite long; abdomen oxypygous with sometimes considerably projecting sheath of ovipositor. First tergite long and thin, postpetiolus only slightly wider than petiolus, without middle field, more or less densely punctured, gastrocoeli in form of elongated longitudinal impressions, interrupted by very large, broadened thyridia; second tergite always considerably longer than width at apex; lunulae absent; at two species from Madagascar base of third tergite with very strong lateral impressions; sculpture of front tergites varies from very densely punctured and almost matt to almost coriaceous.

C o l o r a t i o n: Considerably vary from mainly yellow, ferruginous to red and black with white spots.