Horaeomorphus verus , Jałoszyński, Paweł, 2014

Jałoszyński, Paweł, 2014, ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 3828 (1), pp. 1-76: 17-18

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Horaeomorphus verus

sp. n.

Horaeomorphus verus  sp. n.

( Figs. 26View FIGURES 22 – 28, 31–32View FIGURES 29 – 32, 35– 36View FIGURES 33 – 36, 193View FIGURES 193 – 198 b)

Type material studied. Holotype: AUSTRALIA: ♂: three labels: " 16.05 S 145.28 E QLD / CT 2 10m, 2 km SSW / Cape Tribulation / 1-28 Feb 1996 / L.Umback / FI Trap JCU" [white, printed], " ANIC / Specimen" [green, printed], " HORAEOMORPHUS  / verus  m. /, det. P. Jałoszyński, 2013 / HOLOTYPUS " [red, printed] ( ANIC).

Diagnosis. Male: BL> 2 mm; pronotum with three antebasal pits connected by distinct transverse groove. Male: aedeagus in ventral view with apical part broadly rounded.

Description. Body ( Fig. 26View FIGURES 22 – 28) flattened, elongate and slender, with moderately long appendages, BL 2.05 mm; cuticle glossy, body uniformly dark brown with slightly lighter appendages, vestiture light brown.

Head ( Fig. 26View FIGURES 22 – 28) broadest at eyes, HL 0.33 mm, HW 0.41 mm; tempora slightly longer than eyes, strongly curving posteromesally; vertex strongly transverse, more convex on sides than in middle, with distinctly concave posterior margin; frons subtrapezoidal, posteriorly confluent with vertex, anteriorly gradually lowering toward clypeus; supraantennal tubercles prominent, each sharply demarcated from median part of frons and indistinctly demarcated laterally from vertex, each accompanied by pair of deep pits near posteromesal margin. Eyes moderately large, bean-shaped, weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, inconspicuous; setae short, sparse and suberect, except for sparse and short erect bristles on tempora. Antennae ( Fig. 26View FIGURES 22 – 28) slender, AnL 0.88 mm; antennomeres I –VI each distinctly elongate, VII about as long as broad, VIII –X strongly transverse, XI shorter than IX –X together, about 1.5 x as long as broad, with indistinctly pointed apex.

Pronotum ( Fig. 26View FIGURES 22 – 28) distinctly elongate and weakly convex, broadest near anterior third, PL 0.58 mm, PW 0.53 mm; anterior and lateral margins rounded; sides moderately strongly convergent posteriorly; hind pronotal corners distinct but obtuse and blunt; posterior margin weakly arcuate; base of pronotum with three small but deep and distinct pits connected by sharply marked transverse groove. Punctures on pronotal disc small but distinct, in middle of disc separated by spaces about as wide as 1-1.5 x diameters of punctures; setae moderately dense, thin, short and suberect, sides of pronotum with bristles.

Elytra ( Fig. 26View FIGURES 22 – 28) oval, as convex as pronotum, broadest slightly behind anterior third, EL 1.15 mm, EW 0.75 mm, EI 1.53; basal impressions short; humeral calli developed as elongate protuberances distinctly demarcated mesally from basal part of elytra; elytra with indistinct circumsutural impression near anterior third, small area behind impression slightly raised; elytral apices rounded together. Punctures on elytra unevenly distributed, those on circumsutural impression larger and deeper than those on pronotum, sharply marked and separated by spaces as wide as 0.5–1 diameters of punctures, punctures reducing in diameter, depth and sparser toward elytral margins; setae moderately long and dense, suberect. Hind wings well developed, about twice as long as elytra.

Legs ( Fig. 26View FIGURES 22 – 28) moderately long and slender, without modifications.

Aedeagus ( Figs. 31 –32View FIGURES 29 – 32, 35– 36View FIGURES 33 – 36) moderately stout, AeL 0.40 mm, in ventral view apical part subtrapezoidal, apical margin rounded in middle; internal armature darkly sclerotized and complicated, symmetrical, with long subapical median tubular structure and bell-shaped central complex of sclerites; parameres slender, not exceeding apex of median lobe, with long apical setae.

Female. Unknown.

Etymology. The Latin adjective verus  (real, true, genuine) was chosen as a specific epithet, to underline that this is the first and only true Australian Horaeomorphus  originally placed in a correct genus.

Distribution ( Fig. 193View FIGURES 193 – 198 b). North-eastern Australia (northern Queensland).

Remarks. Horaeomorphus verus  has the aedeagus similar to that of H. anthicoides  , but clearly differs in the external structures; e.g., the body length, proportions of body parts, the sculpture of pronotal base and much longer and denser setae on the dorsum. For characters separating this species from Oriental congeners see the Remarks section at the redescription of H. anthicoides  .


Australian National Insect Collection