Scydmaenozila mallacootana , Jałoszyński, Paweł, 2014

Jałoszyński, Paweł, 2014, ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 3828 (1), pp. 1-76: 37

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Scydmaenozila mallacootana

sp. n.

Scydmaenozila mallacootana  sp. n.

( Figs. 74View FIGURES 72 – 79, 85–86View FIGURES 80 – 88, 93– 94View FIGURES 89 – 96, 195View FIGURES 193 – 198 c)

Type material studied. Holotype: AUSTRALIA: ♂: four labels: " AUSTRALIA / Victoria / Mallacoota N.P. / 26 May 1978 / S. & J. Peck" [white, printed], "mushroom / litter' [white, printed], "Aust. Nat. / Ins. Coll." [green, printed], " SCYDMAENOZILA  / mallacootana  m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] ( ANIC).

Diagnosis. Male and female: body uniformly brown. Male: aedeagus slender, in ventral view with broadly rounded apex; apical part of median lobe abruptly demarcated.

Description. Body of male ( Fig. 74View FIGURES 72 – 79) strongly convex, elongate and slender, with long appendages, BL 2.75 mm; cuticle glossy, dark brown with reddish hue, elytra slightly lighter than head and pronotum, vestiture dark brown.

Head ( Fig. 74View FIGURES 72 – 79) broadest at eyes, HL 0.50 mm, HW 0.60 mm; tempora longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes large and strongly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine, separated by spaces 1.5– 2 x as wide as diameters of punctures; setae on frons and anterior portion of vertex short, sparse and suberect, tempora and posterior portion of vertex densely covered with long bristles directed posteriorly and posterolaterally. Antennae ( Fig. 74View FIGURES 72 – 79) slender, AnL 1.65; antennomeres I –VII distinctly elongate, VIII slightly longer than wide, IX –X each about as long as broad, XI as long as IX –X together, 2.5 x as long as broad, pointed.

Pronotum ( Fig. 74View FIGURES 72 – 79) in dorsal view strongly elongate, broadest near anterior third, PL 0.80 mm, PW 0.63 mm; anterior and lateral margins in anterior half rounded; sides slightly concave in posterior third; posterior margin distinctly bisinuate; base of pronotum with two pairs of large and deep pits, additionally with a pair of shallow but distinct impressions located dorsolaterally. Punctures on pronotal disc fine and inconspicuous; entire dorsal and lateral surface of pronotum covered with moderately dense bristles, dorsally with admixture of thin setae.

Elytra ( Fig. 74View FIGURES 72 – 79) slightly rhomboidal, broadest near middle, EL 1.45 mm, EW 1.05 mm, EI 1.38; basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2– 3 x as wide as diameters of punctures; setae long, curved, sparse and erect. Hind wings well developed, about twice as long as elytra.

Legs ( Fig. 74View FIGURES 72 – 79) long and slender, without modifications.

Aedeagus ( Figs. 85 –86View FIGURES 80 – 88, 93– 94View FIGURES 89 – 96) elongate and relatively slender; AeL 0.63 mm; in ventral view short apical part abruptly demarcated, with broadly and weakly rounded apical margin; in lateral view apical part subtriangular and sharply demarcated from ventral wall; internal armature complicated, with submedian complex of elongate and darkly sclerotized structures and subapical finely denticulate lateral lobes of internal sac; parameres long and slender, with distinctly broadened apices.

Female. Unknown.

Distribution ( Fig. 195View FIGURES 193 – 198 c). South-eastern Australia: Victoria.

Etymology. Locotypical, after the Mallacoota National Park, where this species occurs.

Remarks. Scydmaenozila mallacootana  has the aedeagus very similar to that of Scydmaenozila macrosticta  ; differences can be seen only in the arrangement of internal structures. However, externally these species are distinctly different: Scydmaenozila macrosticta  has the bicolorous pattern on elytra, while Scydmaenozila mallacootana  is uniformly brown; also the shape of the pronotum and elytra is different ( Fig. 72View FIGURES 72 – 79 vs. 74).


Australian National Insect Collection