Scydmaenozila inexpectata, Jałoszyński, Paweł, 2014

Jałoszyński, Paweł, 2014, ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 3828 (1), pp. 1-76 : 37-38

publication ID	https://doi.org/ 10.11646/zootaxa.3828.1.1
publication LSID	lsid:zoobank.org:pub:B2FEAE60-7E51-45FA-A38F-930A084A5AAA
DOI	https://doi.org/10.5281/zenodo.6141074
persistent identifier	https://treatment.plazi.org/id/03F68791-FFF3-182D-8FFE-3C890677F931
treatment provided by	Plazi
scientific name	Scydmaenozila inexpectata
status	sp. nov.

Treatment

Scydmaenozila inexpectata View in CoL sp. n.

( Figs. 75 View FIGURES 72 – 79 , 87–88 View FIGURES 80 – 88 , 95–96 View FIGURES 89 – 96 , 195 View FIGURES 193 – 198 d)

Type material studied. Holotype: AUSTRALIA: ♂: five labels: "Blue Mts. N. S. Wales " [white, printed], "F. W. Ferguson / Collection" [white, printed], " Phagonophana / latipennis Lea / N. S. Wales " [white, handwritten], "Aust. Nat. / Ins. Coll." [green, printed], " SCYDMAENOZILA / inexpectata m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] ( ANIC). Paratype: 1 ♀, originally mounted on one card together with the holotype, labeled with white, printed copies of original labels, and with standard yellow " PARATYPUS " label ( ANIC).

Diagnosis. Male and female: body uniformly brown. Male: aedeagus slender, in ventral view with rounded and subtriangular apex; apical part of median lobe not demarcated.

Description. Body of male ( Fig. 75 View FIGURES 72 – 79 ) strongly convex, elongate and slender, with long appendages, BL 2.98 mm; cuticle glossy, uniformly reddish-brown, vestiture slightly lighter than cuticle.

Head ( Fig. 75 View FIGURES 72 – 79 ) broadest at eyes, HL 0.50 mm, HW 0.60 mm; tempora longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes large and strongly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine, separated by spaces 1.5– 2 x as wide as diameters of punctures; setae on frons and anterior portion of vertex short, sparse and suberect, tempora and posterior portion of vertex densely covered with long bristles directed posteriorly and posterolaterally. Antennae ( Fig. 75 View FIGURES 72 – 79 ) slender, AnL 1.78; antennomeres I–VIII distinctly elongate, IX–X each about as long as broad, XI slightly shorter than IX–X together, slightly less than twice as long as broad, pointed.

Pronotum ( Fig. 75 View FIGURES 72 – 79 ) in dorsal view strongly elongate, broadest near anterior third, PL 0.88 mm, PW 68 mm; anterior and lateral margins in anterior half rounded; sides slightly concave in posterior third; posterior margin distinctly bisinuate; base of pronotum with two pairs of large and deep pits, additionally with a pair of shallow but distinct impressions located dorsolaterally. Punctures on pronotal disc fine and inconspicuous; entire dorsal and lateral surface of pronotum covered with moderately dense bristles, dorsally with admixture of thin setae.

Elytra ( Fig. 75 View FIGURES 72 – 79 ) regularly oval, broadest near middle, EL 1.60 mm, EW 1.10 mm, EI 1.45; basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2– 3 x as wide as diameters of punctures; setae long, curved, sparse and erect. Hind wings well developed, about twice as long as elytra.

Legs ( Fig. 75 View FIGURES 72 – 79 ) long and slender, without modifications.

Aedeagus ( Figs. 87–88 View FIGURES 80 – 88 , 95–96 View FIGURES 89 – 96 ) elongate and slender; AeL 0.60 mm; in ventral view apex not demarcated and subtriangular, with narrowly rounded tip; in lateral view apical part subtriangular and not demarcated from ventral wall; internal armature complicated, with submedian complex of elongate and darkly sclerotized structures and subapical finely denticulate lateral lobes of internal sac; parameres long and slender, with strongly broadened apices.

Female. Slightly larger than male and with shorter antennae, but does not differ in proportions of antennomeres. BL 3.18; HL 0.53 mm, HW 0.63 mm, AnL 1.65 mm; PL 0.95 mm, PW 0.73 mm; EL 1.70 mm, EW 1.25 mm, EI 1.36.

Distribution ( Fig. 195 View FIGURES 193 – 198 d). South-eastern Australia: New South Wales.

Etymology. The epithet inexpectata refers to the surprising finding of this new species among specimens identified previously by Lea as Phagonophana latipennis .

Remarks. Scydmaenozila inexpectata , although externally similar to the remaining uniformly brown species, differs clearly in the aedeagus, which has the apex subtriangular, and not emarginated or broadly rounded.