Spinosciacharis clydensis , Jałoszyński, Paweł, 2014

Jałoszyński, Paweł, 2014, ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 3828 (1), pp. 1-76: 48-49

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Spinosciacharis clydensis

sp. n.

Spinosciacharis clydensis  sp. n.

( Figs. 115View FIGURES 112 – 116, 119View FIGURES 117 – 120, 125– 126View FIGURES 121 – 128, 196View FIGURES 193 – 198 c)

Type material studied. Holotype: AUSTRALIA: ♂: four labels: " AUSTRALIA: NSW / Clyde Mt. 18 km / SE Braidwood / 500 m 28 May 78 / S. & J. Peck" [white, printed], "wet sclerophyll / ravines, log / litter and fungi" [white, printed], "Aust. Nat. / Ins. Coll." [green, printed], " Spinosciacharis  / clydensis  m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] ( ANIC).

Diagnosis. Male: BL 1.90 mm; elytra oval; EI <1.5; metafemur with distinctly curved and broadly rounded ventral projection and with distinct distal ventral lobe. Female and its diagnostic characters unknown.

Description. Body ( Fig. 115View FIGURES 112 – 116) flattened, elongate and slender, with long appendages, BL 1.90 mm; cuticle glossy, body light brown, with slightly lighter appendages, vestiture yellowish.

Head ( Fig. 115View FIGURES 112 – 116) rounded, broadest just behind eyes, HL 0.38 mm, HW 0.40 mm; tempora nearly three times as long as eyes, strongly curved posteromesally; vertex evenly convex; frons confluent with vertex, subtrapezoidal, convex. Eyes small and weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum inconspicuous, fine and sparse, not larger than those on pronotum; setae on frons and median part of vertex short, sparse and suberect, tempora and sides of vertex with moderately dense bristles directed posterolaterally. Antennae ( Fig. 115View FIGURES 112 – 116) slender, AnL 0.85 mm; antennomeres I –VI distinctly elongate, VII –VIII about as long as broad, IX strongly and X slightly transverse, XI much shorter than IX –X together, 1.6 x as long as broad, pointed.

Pronotum ( Fig. 115View FIGURES 112 – 116) in dorsal view elongate, broadest near anterior third, PL 0.50 mm, PW 0.45 mm; anterior and lateral margins rounded; posterior margin indistinctly bisinuate; base of pronotum with distinct but short sublateral carinae accompanied mesally by shallow impressions, median part of pronotal base with two small but deep pits separated by narrow wrinkle as long as pits and not extending beyond pits, but posterior pronotal margin with indistinct median protuberance, so that the median wrinkle between pits can be described as interrupted behind pits. Punctures on pronotal disc fine and inconspicuous, similar to those on frons and vertex; setae sparse, thin, short and suberect, only sides of pronotum with moderately dense bristles and in anterior half with several extremely long, erect setae directed laterally.

Elytra ( Fig. 115View FIGURES 112 – 116) slightly rhomboidal, broadest distinctly anterior to middle, EL 1.03 mm, EW 0.70 mm, EI 1.46; basal impressions short but deep; circumsutural area in anterior fourth barely noticeably more convex than surrounding surface and posteriorly accompanied by distinct and large but shallow impression; elytral apices rounded together. Punctures on elytral disc fine but slightly more distinct and denser than those on pronotum; setae short, sparse and suberect. Hind wings well developed, about twice as long as elytra.

Legs ( Figs. 115View FIGURES 112 – 116, 119View FIGURES 117 – 120) long and slender, metafemur ( Fig. 119View FIGURES 117 – 120) with long and distinctly curved ventral projection slightly broadening toward broadly rounded apex, with distinct ventral distal lobe.

Aedeagus ( Figs. 125–126View FIGURES 121 – 128) in the only known male with completely everted internal armature, AeL (without everted parts) 0.33 mm, in ventral view sides of median lobe slightly concave in subapical region, apical margin broadly rounded; internal armature lightly sclerotized, with two strongly elongate and subtriangular lateral projections protruding distally, in lateral view with a pair of dorsally-directed hook-like projections; parameres distinctly broadening distally and their apices only slightly exceeding apex of median lobe.

Female. Unknown.

Etymology. Locotypical, after the Clyde Mountain.

Distribution ( Fig. 196View FIGURES 193 – 198 c). South-eastern Australia: New South Wales.

Remarks. Spinosciacharis clydensis  resembles Spinosciacharis acheron  sp. n. described below; both species are small and have a similar body silhouette and proportions of body parts, also both have several extremely long lateral setae on sides of pronotum. The shape of metafemoral projection is clearly different ( Fig. 119View FIGURES 117 – 120 vs. 120): curved and slightly broadening distally in Spinosciacharis clydensis  vs. recurved and narrowing distally in Spinosciacharis acheron  . Moreover, the antennae in Spinosciacharis clydensis  are stouter in their distal half, the antennomere VII is nearly as long as broad, while in Spinosciacharis acheron  the antennomere VII is distinctly elongate; also the antebasal median longitudinal carina or wrinkle separating internal pair of pits in Spinosciacharis clydensis  is interrupted behind pits, while in Spinosciacharis acheron  it is continuous.


Royal Botanic Gardens, National Herbarium of New South Wales


Australian National Insect Collection