Clistoabdominalis uzbekistanus ( Kozánek, 1988 ) Kehlmaier, 2005

Kehlmaier, Christian, 2005, Taxonomic studies on Palaearctic and Oriental Eudorylini (Diptera: Pipunculidae), with the description of three new species, Zootaxa 1030 (1), pp. 1-48 : 33-35

Clistoabdominalis uzbekistanus ( Kozánek, 1988) comb. nov. ( Figs 11a–g )

Eudorylas uzbekistanus Kozánek, 1988: 945 .

Material studied

Uzbekistan 1♂ (holotype), Uzbekistan , Kizil­kir (fields), 50km NW of Buchara, 11.VI.1982, leg. M. Chvála, coll. SNMB (Type No. 569) .

Redescription

Male

Body length. About 3.5mm.

Head. Face dark, silver­grey pollinose. Scape dark, without upper bristle. Pedicel with two to three short upper and two dark short lower bristles. Flagellum missing. Eyes meeting for eleven times diameter of ocellus. F:EM:V=1.0:0.6:0.8 (estimated). Frons dark, silver­grey pollinose. Vertex dark, lacking pollinosity, bearing an equilateral, slightly elevated ocellar triangle. Occiput dark, weakly grey pollinose, changing to brown in upper fifth.

Thorax. Pleura, prescutum, scutum and scutellum dark. Pleura grey pollinose. Postpronotal lobe pale, grey pollinose, with about four basal postpronotal hairs present. Prescutum and scutum grey pollinose along anterior fifth, laterally down to wing base and narrowly along posterior margin incl. postalar callus. Otherwise brown pollinose, with two uniseriate dorsocentral rows and some supra­alar hairs. Scutellum brown pollinose, with a fringe of about eight short hairs.

Wing. Length: About 4.3mm. LW:MWW=about 3.2. Wing with microtrichia absent or reduced in small basal cells of wing, e.g. bc, and the beginning of cells c, r 1, br, bm, cup and anal lobe. In cell sc, the microtrichia are partly reduced in the middle part. Pterostigma complete (LS:LTC=1.0). LTC:LFC=1.0 (estimated). r­m reaches dm between basal quarter and third of the cells length. M 1 gently undulating in middle part.

Halter . Base and knob dark, stem white.

Legs. Coxae dark, grey pollinose. Front and mid coxae with pale anteroapical margins. Mid coxa with three to four anterior bristles on inner apical corner. Trochanters pale, partly grey pollinose. Front femur pale but dorsally somewhat darkened, grey pollinose. Mid femur missing. Hind femur dark, with pale base and apex, grey pollinose but shining posteroventrally. Front femur with some dark antero­ and posteroventral peg­like spines towards apex. Front and hind tibiae pale, grey pollinose. Front tibia with small distal spines. Hind tibia with a wrinkled indentation midanteriorly and bent by 135° (inner angle) at its middle. Tarsi pale with distitarsi darkened, weakly grey pollinose. Pulvilli slightly shorter than distitarsi.

Abdomen. Ground colour dark. Tergite 1 with three dark lateral bristles. Tergite 1 anterodorsally brown, otherwise grey pollinose. Tergite 2 to 5 (antero­) laterally grey pollinose, dorsally brownish­bronze pollinose. Viewed dorsally, tergite 3 to 5 very narrow. Sternite 1 to 5 dark, greyish pollinose. Sternite 6 dark, greyish pollinose. Sternite 7 dark. Syntergosternite 8 dark and enlarged, without a distinct membranous area but with a weaker sclerotized area in its middle. Viewed laterally, abdomen with a large and conspicuous ventral projection, its ventral part being made up by sternite 5 (with many bristly hairs), its posterior inner part being predominantly made up by sternite 6 and 7. Syntergosternite 8 directed upwards.

Genitalia.

Genital capsule dorsal view: Epandrium paler than tergites and longer than wide (MLE:MWE=1.4). Surstyli paler than tergites and rather symmetrical ( Fig. 11d ). Both are of triangular ground shape with a tapering apical projection.

Genital capsule ventral view: Gonopods subequal, inner one minute, outer one small but clearly larger ( Fig. 11a ). Phallus short and trifid, semicircular with broad ejaculatory ducts ( Fig. 11c ). Phallic guide rather simple: small, broad, straight and cone­shaped ( Fig. 11a ).

Genital capsule lateral view: Epandrium without projecting lobe on either side. Surstyli straight ( Figs 11f–g ). Phallic guide very deep, but rather translucent, and with a dorsal projection close to its apex ( Fig. 11b ).

Ejaculatory apodeme very large, balloon­shaped ( Fig. 11c ).

Female unknown.

Discussion

Due to the shape of the male genitalia, the taxon is hereby transferred to the genus Clistoabdominalis . With its conspicuous ventral projection, illustrated in Kozánek (1988), Cli. uzbekistanus ( Kozánek, 1988) comb. nov. can be readily distinguished from all other known species of this genus. The structure of the inner male genitalia is figured here for the first time. Judging from it, the taxon could be regarded as an intermediate between the electus and ruralis species groups (see Kehlmaier 2005). Cli. uzbekistanus is currently only known from its locus typicus in Uzbekistan.

Figures

FIGURE 11. Clistoabdominalis uzbekistanus (Kozánek, 1988). (a) Phallic guide, gonopods and hypandrium ventral view. (b) Phallic guide lateral view. (c) Phallus ventral view. (d) Surstyli strictly dorsal view. (e) Ejaculatory apodeme. (f) Outer surstylus lateral view. (g) Inner surstylus lateral view. Scale bars = 0.1 mm.