Hermanella costalis (Navás) Navas, 2005

Peters, J. G., Flowers, R. W., Hubbard, M. D., Domínguez, E. & Savage, H. M., 2005, New records and combinations for Neotropical Leptophlebiidae (Ephemeroptera), Zootaxa 1054, pp. 51-60 : 55

Hermanella costalis (Navás) comb. n.

( Figures 10–15 )

Thraulus costalis Navás, 1934:26 ; Traver, 1960:73.

Thraulus costalis Navás, 1934 , was described from Brazil (Barão Homem de Mello, Rio de Janeiro) from a male imago in poor condition. Traver (1960) stated that it was not Thraulus , but she did not reassign the species. Photographs of the holotype were prepared by C. Kutzscher of the Deutsches Entomologisches Institut im ZALF; P. Grant, who examined the holotype in 1985, provided additional notes on its genitalia. The wings ( Fig. 10– 11 ) and genitalia ( Fig. 14–15 ) clearly place the species in the Hermanella generic complex as defined by Domínguez & Flowers (1989), but the condition of the styliger plate (blunt median protuberance and no paired dorsal projections) does not fit the definition of any described genus of the complex. Also, the ventral projection on each penis lobe is broad in Hermanella and slender in T. costalis . Although T. costalis is not a species of Hermanella in the restricted sense, we prefer not to establish a new genus until the nymphs are known, and thus treat this species as Hermanella costalis (Navás) new combination. A set of figures of the imago is included for purposes of future identification ( Fig. 10­15 ). Note that the torn hind wing is hyaline and that the color in Fig. 11 is that of the body showing behind the wing. The only hind wing color is a brown basal strip in the costal area ( Fig. 12 ).

Figures

FIGURE 10 – 15, holotype of Thraulus costalis: 10, fore wing; 11, hind wing enlarged (actual size about 1.5 mm); 12, lateral view of body (with left foreleg); 13, detail of foretarsal segments (outer surface); 14 – 15, male genitalia with detail. Photographs by C. Kutzscher. (Editing: Fig. 12, pin removed; Fig. 11, 14 – 15 details outlined.)