Mada hirsuta, Szawaryn, Karol & González, Guillermo, 2017

Szawaryn, Karol & González, Guillermo, 2017, A new species of Mada Gordon from the Andes (Coleoptera: Coccinellidae: Epilachnini), Zootaxa 4226 (3), pp. 442-446 : 442-445

publication ID

https://doi.org/ 10.11646/zootaxa.4226.3.7

publication LSID

lsid:zoobank.org:pub:A769D6E3-83C0-4BCC-A304-5FC092EF6785

DOI

https://doi.org/10.5281/zenodo.6004530

persistent identifier

https://treatment.plazi.org/id/03F9CF7C-FFB6-DB7B-FF41-F88AFDE6FF0A

treatment provided by

Plazi

scientific name

Mada hirsuta
status

sp. nov.

Mada hirsuta sp. nov.

( Figs. 1– 3 View FIGURE 1 View FIGURE 2 View FIGURE 3 )

Etymology. The species name is derived from its distinctive elytral pubescence. Diagnosis. This species can be quite easily distiguished from other Mada species by the surface of elytra covered with long setae irregularly arranged in whirling pattern ( Fig. 1 View FIGURE 1 A). Moreover it has protibiae distinctively broadened ( Fig. 2 View FIGURE 2 G), male genitalia with penis guide narrow, pointed, shorter than parameres, and female genitalia with coxites much longer than wide, narrow at base and broadened at posterior part.

Description. Length 3.9–4.3 mm; width 3.4–3.7 mm; TL/EW = 1.14–1.16; PL/PW = 0.50–0.51; PL/EL = 0.30– 0.31; PW/EW = 0.57–0.61; EL/EW = 0.95–1.00.

Body oval, convex ( Figs. 1 View FIGURE 1 A, 1B); pronotum and elytra covered with long setae. Head and mouthparts yellowishbrown ( Fig. 1 View FIGURE 1 C); pronotum yellowish-brown with anterior angles more pale, disc with five dark spots, one at base and four in a curved row across middle ( Fig. 1 View FIGURE 1 C); elytra yellowish-brown with lateral margin pale yellow ( Fig. 1 View FIGURE 1 B). Ventral surface light-brown. Head and pronotum punctate; elytron dually punctate.

Head exposed, transverse; ventral antennal grooves present. Eye finely faceted, with inner orbits emarginate anteromedially ( Fig. 1 View FIGURE 1 C). Antenna shorter than width of head, 11-segmented ( Fig. 2 View FIGURE 2 D); scape large, swollen, about twice as long as pedicel; pedicel longer than wide, swollen; antennomere 3 elongate, shorter than antennomeres 4–6 combined; antennomeres 4–7 elongate, antennomere 8 transverse; antennal club relatively compact, 3-segmented, asymmetrical. Clypeus transverse; labrum transverse with anterior part membranous, covered with long setae ( Fig. 2 View FIGURE 2 F). Mandibles multidentate, with large teeth on inner edge, with several small denticles on dorsal surface ( Fig. 2 View FIGURE 2 E).

Pronotum transverse, widest at base and gradually narrowing anteriorly; anterior angles and base not bordered; disc convex, finely punctate ( Fig. 1 View FIGURE 1 C). Prothoracic hypomeron smooth; notosternal suture distinct; prosternal process with weak grooves along its lateral margins. Mesoventrite with anterior edge emarginate with raised border for entire length; mesoventral process smooth, as broad as coxal diameter; meso-metaventral articulation with suture straight. Scutellar shield small, triangular. Elytra at base broader than pronotum; with dual punctation; humeral angles well developed; lateral margins narrow, visible from above throughout ( Fig. 1 View FIGURE 1 A); elytral epipleuron complete, with distinct depressions for receiving apices of mid and hind femora, with submarginal carina complete; metaventrite with intercoxal process broadly bordered; metaventral postcoxal lines forming straight line with two rounded, setose projections, laterally complete, recurved; metaventrite with discrimen complete.

Legs short and stout, not protruding from outer margin of elytral epipleuron. Trochanters roundly produced, with cavities on their inner surfaces for receiving tip of tibiae in repose. Femora cylindrical with weak grooves throughout for receiving tibiae. Tibiae with apical spurs of formula 1-2-2, protibiae expanded with weak grooves throughout for receiving tarsi ( Fig. 2 View FIGURE 2 G), mid and hind tibiae on outer edge near apex with oblique carina; tarsal claw double with additional large, subtriangular basal tooth ( Fig. 2 View FIGURE 2 N).

Abdomen with five ventrites in males with tip of 6th ventrite visible ( Fig. 2 View FIGURE 2 B) and five ventrites in females ( Fig. 2 View FIGURE 2 A). Abdominal postcoxal lines recurved roundly and incomplete, almost reaching posterior margin of ventrite 1 ( Figs. 2 View FIGURE 2 A, B). In males apical margin of ventrite 5 truncate ( Fig. 2 View FIGURE 2 B); ventrite 6 emarginate medially; tergite VIII rounded. Apodeme of male sternum IX simple, thin, rod-like. Tergite X rounded at apex ( Fig. 2 View FIGURE 2 M). In females apical margin of ventrite 5 rounded ( Fig. 2 View FIGURE 2 A); sternite VIII rounded, entire, with membranous part in middle appearing divided into two parts ( Fig. 2 View FIGURE 2 C), tergite VIII rounded. Proctiger (TX) membranous at base, elongate, rounded apically ( Fig. 2 View FIGURE 2 L).

Male terminalia and genitalia ( Figs. 2 View FIGURE 2 H–J). Penis guide symmetrical, thin, with sides almost parallel, spindle shaped, widest at basal part in lateral view, with pointed apex; shorter than parameres. Parameres well developed, simple, setose apically. Penis base with capsule reduced. Penis simple, rod-like.

Female genitalia ( Figs. 2 View FIGURE 2 K, L). Coxites elongate, narrow at base, broadly widened at posterior part, styli not observed. Bursa copulatrix ending with oviduct, sperm duct diverges slightly below oviduct; sperm duct short, lightly sclerotized, simple; spermatheca small, membranous, vermiform, without clear nodulus and ramus; accessory gland thin, a bit longer than spermatheca.

Type material. Holotype, male, Ecuador (Orellana), Payamino Research Station 0°29'36.01''S 77°17'29.15''W 300m Tropical Rainforest Malaise trap, 30.VII-12.VII.2007 / coll GoogleMaps . CPDT Gillet BMNH (E) 2007-65 ( BMNH). Paratypes, ECUADOR: 1 female , Ecuador (Orellana), Payamino Research Station 0° 29' 36.01''S 77° 17' 29.15''W 300m Tropical Rainforest, malaise trap, 30.VII–12.VII.2007 / coll GoogleMaps . CPDT Gillet BMNH (E) 2007-65 ( BMNH) ; PERU: 1 female , Peru, Madre de Dios, CICRA Field Stn , garden. 12.56940°S 70.10100°W 260m, 25.X–1.XI.2010, MJ Endara, malaise trap GoogleMaps , PER10-10-MAT-021/ SEMC1097096 KUNHM-ENT (MUSM).

Distribution. Ecuador, Peru ( Fig. 3 View FIGURE 3 ).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Coccinellidae

Genus

Mada

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