Habronestes macedonensis ( Hogg, 1900 )

Baehr, Barbara, 2003, Revision of the Australian Spider Genus Habronestes (Araneae: Zodariidae). Species of New South Wales and the Australian Capital Territory, Records of the Australian Museum 55 (3), pp. 343-376 : 351-352

Habronestes macedonensis ( Hogg, 1900)

Figs. 35, 36 , 37 , 40, 41 , 46 , 139

Storena macedonensis Hogg, 1900: 2 , 97, 98, pl. 14, fig. 4 (description male); Rainbow, 1911:150 (description male).

Habronestes macedonensis .– Jocqué, 1995: 145, fig. 3c,d (description male).

Remarks. The male of Habronestes macedonensis was first described by Hogg, 1900 as Storena macedonensis . Jocqué (1995) has reexamined the holotype and placed the species in Habronestes because of the special palp structure. The female is described here for the first time.

Material examined. 1♀, 13: NSW, Washpool NP Grassy Ck, where crossed by North West Fire Trail / Washpool Trail, 29°27'43"S 152°16'22"E, 4.ii–9.iv. 1993, 900 m, G/C, AM KS36298 ; 1 ♀, 13 as previous, QM S60828 ; 13 Conglomerate SF, 100 m SW of jnctn of Hallgraths Trail & Sherwood Rd, 30°7'0"S 153°3'14"E, 4.ii–9.iv. 1993, 320 m, G/C, AM KS39028 ; 13 Hazelbrook , 33°44'S 150°27'E, 4.ii.1979, M. Dingley, AM KS3037 ; 13 Warra SF, 2.8 km W of Moggs Swamp Ck, Moggs Swamp Fire Trail , 29°59'19"S 151°57'14"E, 4.ii–9.iv.1993, 1140 m, G/C, AM KS36361 ; 13 Warra SF, Moggs Swamp Fire Trail, 2.5 km N of Moggs Swamp Ck, 29°57'32"S 151°58'18"E, 4.ii–9.iv.1993, 1170 m, G/C, AM KS36370 ; 13 Rocky Waterholes Rd , Beaury SF, 28°32'49"S 152°20'11"E, 9.iv.1993, 4.ii–9.iv. 1993, 705 m, G/C, AM KS56172 . ACT: 1 3 Tidbinbilla SF, 35°26'S 148°56'E, litter, P. Ormay, AM KS3535 .

Diagnosis. Eyes in 2 rows, both rows procurved, AME largest. Abdomen with 2 pairs of white patches on anterior part, the first elongated and nearly divided in two; and 2 patches in front of spinnerets. Metatarsus I with irregular band of short strong spines ventrally in males. LTA fanshaped covered with small spicules ( Figs. 40, 41 ). Epigyne with w-shaped scape. Vulva with extremely short copulatory ducts ending in large, globular, not touching spermathecae ( Figs. 35, 36 ).

Description. Male (AM KS36298). Total length 6.24; carapace 3.44 long, 2.24 wide; 1.32 high; cl/cw 1.53; sternum 1.40 long, 1.36 wide; sl/sw 1.02. Abdomen 2.8 long, 2.16 wide. Colour. Carapace chestnut brown with dark fovea and dark bifurcate patches in front; sternum yellow brown; chelicerae medium brown; maxillae and labium pale brown, distally white. Abdomen sepia brown with 2 pairs of white patches on anterior part, the first elongated and nearly divided in two and 1 long and undulate patch in front of spinnerets; laterally sepia brown with 2–3 long and broad white patches; ventrally pale pink brown. Legs yellow; clearly annulated; legs I–IV with white coxa, prolateral suture sepia brown, pale trochanter, proximal ½ of femora white and distal ½ sepia brown with darker brown lateral stripes on patella and tibiae III, IV. Eyes. In 2 rows each with 4 eyes, both rows procurved. AME largest. Eye group width 0.6 of head width; AME 0.24; ALE 0.16; PME 0.16; PLE 0.16; AME–AME 0.04; AME–ALE 0.04; ALE–PLE 0.04; PME–PME 0.16; PME–PLE 0.16. MOQ. AME–PME 0.52; AME–AME 0.52; PME–PME 0.48. Clypeus & chilum. Clypeus 0.60 high; chilum undivided, short. Legs. Length formula 4132; metatarsus I with irregular band of short, strong spines ventrally; femora III, IV with a row of 5–6 stout dorsal spines at distal end; patella III, IV with 1 irregular longitudinal row of stout spines prolaterally. Male palp ( Figs. 40, 41 , 46 ). Cymbium with flat RCF, ½ of the cymbium length. LTA with a thick, turned vertical stalk and with a sickle-shaped plate serrated on retrolateral margin; DTA chitinous, with a flattened stalk and a curled apical end covered with spicules; VTA sickle-shaped, long, with rounded tip. Tibiae short; with 1 long prolateral seta and with a group of bent spines retrolaterally; DTiA with chitinous rim, triangular; VTiA short and triangular.

Female (AM KS36298). Total length 8.72; carapace 4.32 long, 3.00 wide; 2.00 high; cl/cw 1.50; sternum 1.64 long, 1.60 wide; sl/sw 1.02. Abdomen 4.40 long, 3.20 wide. Colour. Same as male. Eyes. In 2 rows each with 4 eyes, both rows procurved. AME largest. Eye group width 0.54 of head width; AME 0.30; ALE 0.2; PME 0.20; PLE 0.20; AME–AME 0.04; AME–ALE 0.04; ALE–PLE 0.04; PME– PME 0.20; PME–PLE 0.20. MOQ: AME–PME 0.68; AME–

AME 0.64; PME–PME 0.60. Clypeus 1.12 high. Epigyne ( Figs. 35, 36 ). With w-shaped scape. Vulva with extremely short copulatory ducts ending in large globular, not touching spermathecae.

Distribution. Victoria, northeastern New South Wales and ACT ( Fig. 139 ).

Figures

Figs. 31–36. Habronestes macedonensis species-group epigynes, ventral view (above), vulvae, dorsal view (below): (31–32) Habronestes rawlinsonae n.sp.; (33–34) Habronestes hebronae n.sp.; (35–36) Habronestes macedonensis (Hogg). Scales 0.5 mm.

Figs. 37–39. Habronestes macedonensis species-group body, dorsal view. (37) Habronestes macedonensis (Hogg); (38) Habronestes ungari n.sp.; (39) Habronestes weelahensis n.sp. Scales 1 mm.

Figs. 40–45. Habronestes macedonensis species-group male palps, ventral view (above), lateral view (below): (40–

Figs. 46–50. Habronestes macedonensis species-group male palps, LTA, ventral view: (46) Habronestes macedonensis (Hogg); (47) Habronestes weelahensis n.sp.; (48) Habronestes hebronae n.sp.; (49) Habronestes ungari n.sp.; (50) Habronestes rawlinsonae n.sp. Scales 0.25 mm.

Figs. 138–143. New South Wales, showing collection localities: (138) species of Habronestes australiensis species-group; (139) species of Habronestes macedonensis species-group; (140–143) species of Habronestes pictus species-group.