Archaeotinodes petropolitana Melnitsky & Ivanov

Melnitsky, Stanislav I. & Ivanov, Vladimir D., 2013, Three new caddisflies species of the fossil genus Archaeotinodes (Insecta: Trichoptera: Ecnomidae) from the Baltic Amber, Zootaxa 3635 (3), pp. 261-268 : 262-263

Archaeotinodes petropolitana Melnitsky & Ivanov , new species

Figs. 1–2

Description ( Figs. 1–2 ). Abdomen and thorax light brown. Head, antennae and palpi slightly lighter. Wings brown, heavily covered with fine hairs. Anterior branch of CuA in each forewing with additional fork. Third segment of each maxillary palp slightly longer and thicker than second and fourth. Forewings each with five forks. Antennae reaching twothirds of forewing length. Medial hair warts on pronotum large, swollen and united. Scutal hair warts of mesonotum with oval shape and closely approximate. Occipital hair warts of head large, each with rounded posterior margin. Ocellar hair warts narrow and elongated. Anterior margin of abdominal sternite V with distinct, clearly marked cuticular plate, darker than yellow sternites. Small bulge apparently associated with pheromone gland situated postero-medially to this structure. Cuticle of abdominal sternite V covered by small network pattern making rough surface. Spur formula 3.4.4. Measurements: body length 4.7 mm; forewing length 5.3 mm.

Male genitalia ( Fig. 1–2 ). Lower appendages (gonopods) small, subdivided into dorsal and ventral lobes (branches), widely separated. Dorsal lobes oblong and rounded at apex, outer surfaces covered with numerous setae. Ventral lobe of each gonopod long and thin, with apical portion extended into long sickle-shaped and pointed process directed posteroventrad. Aedeagus strong and well sclerotized. Apical part of aedeagus reaching ventral lobes of lower appendages. Sternite IX broad, projecting backwards, with two incisions on ventral margin. Preanal appendages (cerci) very long and curved, enlarged apically; their apical parts without spine. Numerous hairs covering subapical part of outer surface of preanal appendages. The lateral margins of preanal appendages heavily sclerotized. Segment X membranous. Ventral branches of segment X very long, s-shaped and curved.

Comparison. The new species is similar to Archaeotinodes lanceolata Ulmer, 1912 , Archaeotinodes pauper Ulmer, 1912 , and Archaeotinodes igneusaper Melnitsky, 2009 , from Baltic amber. Archaeotinodes petropolitana sp. nov. differs by its longer and more strongly curved apically ventral lobe of the lower appendage, by the more massive aedeagus and the absence of a long black inner spine in the apical part of each preanal appendage (the ventral branch of segment X).

Holotype male. PIN, № 364/789, Baltic amber, Eocene.

Etymology. From the Latin Petropolitana—St. Petersburg. The gender of Archaeotinodes is feminine. Ulmer (1912) selected A. prisca (Pictet) as the type of the genus, and thus, designated the gender of the genus as feminine (See ICZN article 30.1.4.4.) intentionally.

Figures

FIGURE 1. Archaeotinodes petropolitana sp. nov., holotype PIN, № 364 / 789.

FIGURE 2. Archaeotinodes petropolitana sp. nov., holotype PIN, № 364 / 789, male genitalia, lateral. Scale bar = 0.1 mm. Abbreviations: AE—aedeagus, DL—dorsal lobe of lower appendage, PA—preanal appendage, VL—ventral lobe of lower appendage.