Zyras (Zyras) latibasalis, Assing, 2017

Assing, Volker, 2017, On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini), Beiträge Zur Entomologie = Contributions to Entomology 67 (2), pp. 213-246 : 229-230

Zyras (Zyras) latibasalis spec. nov.

urn:lsid:zoobank.org:act:35276B1A-6492-4F55-8836-91058719625C

( Figs 3, 25 , 39–40 , 70–71 )

Type material examined: Holotype ♂: “ INDONESIA: Jawa Barat, Cianjur, Kebun Raya, Cibodas (1400 m alt.), 29.I.–4.II.2004 (FIT), Maruyama M. et al. / Holotypus ♂ Zyras latibasalis sp. n., det. V. Assing 2017” (cMar) . Paratypes: 9 exs.: same data as holotype (cMar, cAss) .

Additional material: 1 ♀: “NE-LAOS: Hu Phan prov. , Ban Saluei, Pu Phan mts., 20°15'N, 104°02'E, 1500–2000 m, 2.IV.–11.V.2001, leg. D. Hauck ” ( NMP) .

Etymology: The specific epithet (with broad base) is an adjective alluding to the basally broad ventral process of the aedeagus in ventral view, the most prominent character distinguishing this species from the similar Z. wei .

Description: Body length 5.5–7.0 mm; length of forebody 2.5–3.0 mm. Coloration ( Figs 3, 25 , 39 ): forebody blackish with the humeral and anterior portions of the elytra sometimes narrowly and diffusely paler; abdomen black with the posterior margins of the tergites more or less distinctly paler reddish; legs pale-yellowish; antennae blackish with antennomeres IX–X often paler brown (X more so than IX) and antennomere XI dark-yellowish to pale-reddish; maxillary palpi pale-brown with the apical palpomere yellowish.

Head ( Fig. 25 ) distinctly transverse; punctation moderately coarse and moderately dense; median dorsal portion extensively impunctate. Eyes large, much longer than postocular region in dorsal view. Antenna ( Fig. 3 ) 1.9–2.2 mm long and moderately massive; antennomeres IV–VI approximately as long as broad, VII–X weakly transverse, X less than 1.5 times as broad as long, and XI approximately as long as the combined length of IX and X.

Pronotum ( Fig. 25 ) strongly transverse, 1.22–1.24 times as broad as long and 1.29–1.36 times as broad as head, broadest near anterior angles; posterior angles weakly, obtusely marked; lateral margins not sinuate in posterior half in dorsal view; punctation moderately coarse and moderately dense, nearly regularly distributed; midline rather narrowly impunctate.

Elytra ( Fig. 25 ) approximately 0.85 times as long as pronotum; punctation rather dense and coarse, slightly denser near scutellum and near suture than in posterior and lateral portions. Hind wings present. Metatarsomere I approximately as long as the combined length of II–IV.

Abdomen ( Fig. 40 ) slightly narrower than elytra, with moderately deep anterior impressions on tergites III–V; tergites III–V each with a transverse row or band of rather fine and often sparse non-setiferous punctures in anterior impressions, with a lateral setiferous puncture on either side and with four (tergite III) or 6–8 (tergites IV–V) setiferous punctures at posterior margins; tergite VI with a narrow transverse band or a transverse row of rather fine non-setiferous punctures anteriorly, with a lateral setiferous puncture on either side, and with usually eight setiferous punctures at posterior margin (individual punctures may be missing, or additional punctures may be present); tergite VII with a transverse band of fine non-setiferous punctures anteriorly and with two transverse rows of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with setiferous punctures bearing long dark setae only in posterior fourth, anteriorly with a transverse band of oblong microsculpture (microstriae), posterior margin with shallow concavity in the middle ( Fig. 39 ).

♂: posterior margin of sternite VIII broadly truncate in the middle; median lobe of aedeagus ( Figs 70–71 ) 0.70–0.75 mm long; ventral process broadly triangular in ventral view; parameres approximately 1.0 mm long, with long and slender apical lobe.

♀: posterior margin of sternite VIII shallowly concave in the middle.

Intraspecific variation: The female from Laos is similar to the type series in size, habitus, coloration, and punctation, except that the apical antennomeres are blackish.

Comparative notes: In external characters (size, habitus, punctation, coloration), in the lateral aspect of the ventral process of the aedeagus, and regarding the long apical lobe of the paramere, Z. latibasalis strongly resembles the widespread Z. wei . It is distinguished from this species (and from the similar Z. parvilobatus ) by the darkyellowish antennomere XI, by the presence of an anterior transverse band of microstriae on tergite VIII, and by the morphology of the aedeagus ( Z. wei : ventral process much more slender and basally somewhat constricted in ventral view, and smoothly curved in lateral view; crista apicalis more prominent).

Distribution: The type locality is situated in Jawa Barat at an altitude of 1400 m. The tentative record from Laos requires confirmation based on males. The type specimens were collected with flight interception traps.

Figures

Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm.

Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm.

Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm.

Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm.