Myrmarachne cyrtodens, Yamasaki, Takeshi & Ahmad, Abdul Hamid, 2013

Yamasaki, Takeshi & Ahmad, Abdul Hamid, 2013, Taxonomic study of the genus Myrmarachne of Borneo (Araneae: Salticidae), Zootaxa 3710 (6), pp. 501-556 : 503-505

Key to species of Myrmarachne of Borneo

Male

1 Pedicel almost as long as, or longer than ALE–PLE ( Figs 6 A, 13A, 32A)........................................ 2

- Pedicel distinctly shorter than ALE–PLE ( Figs 2 A, 4A)...................................................... 4

2 Fang bearing tooth-like apophysis ( Fig. 32 D)...................................... melanocephala MacLeay, 1839

- Fang without tooth-like apophysis....................................................................... 3

3 Chelicera apically swollen............................................................. assimilis Banks, 1930

- Chelicera without swollen apex........................................................ cornuta Badcock, 1918

4 Basal half of fang strongly arched ( Figs 8 C, 16C, 18B)....................................................... 5

- Fang otherwise...................................................................................... 7

5 With carapace in dorsal view, chelicera very short, less than half the length of carapace........ gedongensis Badcock, 1918

- With carapace in dorsal view, chelicera longer, more than half the length of carapace............................... 6

6 In lateral view upper margin of chelicera convex ( Fig. 8 B)............................... biseratensis Badcock, 1918

- In lateral view upper margin of chelicera not convex ( Fig. 16 B)................................... cyrtodens sp. nov.

7 Chelicera apically swollen............................................................................. 8

- Chelicera apically not swollen.......................................................................... 12

8 Apical prolateral tooth of chelicera bifid ( Figs 4 C, 22D)...................................................... 9

- Apical prolateral tooth of chelicera simple............................................................... 10

9 Chelicera without retrolateral teeth...................................................... alticephalon sp. nov.

- Chelicera with retrolateral teeth.......................................................... lambirensis sp. nov.

10 Body length distinctly less than 5 mm; flange of RTA complex ( Figs 20 E–G)....................... hanoii Żabka, 1985

- Body length usually more than 5 mm; RTA weakly sinuous.................................................. 11

11 Prolateral teeth confined to anterior half of chelicera........................................ opaca (Karsch, 1880)

- Prolateral teeth present along whole length of prolateral margin................................ acromegalis sp. nov.

12 Chelicera bearing one or two spurs on its inner margin ( Figs 10 A, 30A)........................................ 13

- Chelicera without spur on its inner margin............................................................... 14

13 One spur at apex of inner margin of chelicera ( Figs 10 A, 10E).................. borneensis Peckham & Peckham, 1907

- Two spurs in anterior part of inner margin of chelicera, more distal spur very large ( Figs 30 A, 30D).................................................................................................. maxillosa (C. L. Koch, 1846)

14 Chelicera distinctly longer than carapace ( Figs 17 A–B, 28A–B, 35A–B)....................................... 15

- Chelicera not longer than carapace..................................................................... 17

15 Abdomen narrow elongate-oval ( Fig. 28 A)..................................... markaha Barrion & Litsinger, 1995

- Abdomen oval..................................................................................... 16

16 Retrolateral cheliceral teeth confined to posterior part of chelicera................................... endoi sp. nov.

- Retrolateral cheliceral teeth confined to anterior part of chelicera.......................... nitidissima (Thorell, 1877)

17 Chelicera distinctly shorter than carapace; body small, less than 5 mm .......................................... 18

- Chelicera not distinctly shorter than carapace; body not small, more than 5 mm .................................. 19

18 Abdomen with two transverse black bands dorsally........................... mariaelenae Edwards & Benjamin, 2009

- Abdomen without such bands.......................................................... brevichelicera sp. nov.

19 Abdomen with posterior black, broadly transverse band......................... shelfordii Peckham & Peckham, 1907

- Abdomen without such a band......................................................................... 20

20 Embolus of palp broadened apically, with a short process at apex ( Fig. 15 E)...................... crassembolus sp. nov.

- Embolus of palp slender, tapering apicad, with pointed apex ( Fig. 24 E)........................................ 21

21 Anterolateral area of cephalic part above coxa I fringed with white hairs; dorsum of thoracic part strongly swollen............................................................................. malayana Edmunds & Prószyński, 2003

- Anterolateral area of cephalic part without fringe of white hairs; dorsum of thoracic part without distinct swelling................................................................................................ sabahna sp. nov.

Female

1 Abdomen with posterior broad, black, transverse band.......................... shelfordii Peckham & Peckham, 1907

- Abdomen without such a band.......................................................................... 2

2 Pedicel almost as long as, or longer than ALE–PLE ( Figs 7 A, 14A, 33A)........................................ 3

- Pedicel distinctly shorter than ALE–PLE ( Fig. 3 A)......................................................... 5

3 Thoracic part distinctly lower than cephalic part, with weakly convex dorsum ( Fig. 7 B).......... assimilis Badcock, 1918

- Thoracic part slightly lower than cephalic part, with distinctly convex dorsum ( Figs 14 C, 33B)...................... 4

4 Copulatory atria oval; lateral pockets present in front of epigastric furrow ( Figs 33 D–H, 34A–C).................................................................................................. melanocephala MacLeay, 1839

- Copulatory atria elongate-oval; small median pocket present in front of epigastric furrow ( Figs 14 D–F).................................................................................................... cornuta Badcock, 1918

5 Anterior part of sclerotised copulatory ducts twisted ( Figs 9 D, 12D)............................................ 6

- Anterior part of sclerotised copulatory ducts not twisted ( Fig. 25 D)............................................ 13

6 Body very small, distinctly less than 5 mm in total length.................................................... 7

- Body larger, more than 5 mm in total length................................................................ 9

7 Anterior part of sclerotised copulatory ducts twisted more than once; lateral pockets present in front of epigastric furrow ( Fig. 21 D)................................................................................ hanoii Żabka, 1985

- Anterior part of sclerotised copulatory ducts twisted once; median pocket present in front of epigastric furrow ( Figs 12 D–F) 8

8 Abdomen with two transeverse black bands dorsally.......................... mariaelenae Edwards & Benjamin, 2009

- Abdomen without such bands.......................................................... brevichelicera sp. nov.

9 Posterior margin of coxa II and anterior margin of coxa III densely fringed with black hairs ( Fig. 23 C)....................................................................................................... lambirensis sp. nov.

- Posterior margin of coxa II and anterior margin of coxa III not fringed with black hairs............................ 10

10 Median pocket of epigyne with anteromedian indentation ( Figs 9 D–F)..................... biseratensis Badcock, 1918

- Median pocket anteriorly rounded ( Figs 3 D–F)............................................................ 11

11 In lateral view dorsum of cephalic part strongly convex ( Fig. 3 B)............................... acromegalis sp. nov.

- In lateral view dorsum of cephalic part almost flat or weakly convex ( Figs 29 B, 38B)............................. 12

12 White lateral marking formed of white hairs present in anterolateral cephalic part above coxae I and III.......................................................................................... markaha Barrion & Litsinger, 1995

- White lateral marking absent in anterolateral part of cephalic part.................................. sabahna sp. nov.

13 Cephalic part remarkably higher (about 2X) than thoracic part ( Fig. 5 B); spermathecae remarkably large ( Figs 5 D–E)......................................................................................... alticephalon sp. nov.

- Cephalic part slightly higher than thoracic part; spermathecae linear, or roundly swollen but not remarkably large....... 14

14 Spermathecae linear and straight ( Figs 19 D–E)........................................ gedongensis Badcock, 1918

- Spermathecae oval or spherical........................................................................ 15

15 Median pocket slender, longer than broad ( Figs 31 D–E)................................ maxillosa (C. L. Koch, 1846)

- Median pocket broad, broader than long, or indistinct ( Figs 25 D–E)............. malayana Edmunds & Prószyński, 2003

Figures

FIGURE 6. M. assimilis, male. A, chelicera, carapace and pedicel, dorsal view; B, pedicel and abdomen, dorsal view; C, chelicera and carapace, lateral view; D, chelicera and fang, ventral view; E, maxillae, labium, sternum and coxae; F, palp, ventral view; G, palp, lateral view; H, palpal tibia, dorsal view. (Scales. A – E: 1 mm; F – H: 0.25 mm)

FIGURE 2. M. acromegalis sp. nov., male. A, body, dorsal view; B, chelicera and carapace, lateral view; C, chelicera and fang, ventral view; D, maxillae, labium, sternum and coxae; E, palp, ventral view; F, palp, dorsal view. (Scales. A – D: 1 mm; FIGURES E – F: 0.25 mm)

FIGURE 32. M. melanocephala, male. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D, chelicera and fang, ventral view; E, palp, ventral view; F, palp, lateral view; G, palpal tibia, dorsal view. (Scales. A – D: 1 mm; E – G: 0.25 mm)

FIGURE 8. M. biseratensis, male. A, body, dorsal view; B, chelicera and carapace, lateral view; C, chelicera and fang, ventral view; D, maxillae, labium, sternum and coxae; E, palp, ventral view; F, palp, lateral view; G, palpal tibia, dorsal view. (Scales. A – D: 1 mm; E – F: 0. 25 mm)

FIGURE 16. M. cyrtodens sp. nov., male. A, body, dorsal view; B, chelicera and carapace, lateral view; C, chelicera and fang, ventral view; D, maxillae, labium, sternum and coxae; E, palp, ventral view; F, palp, lateral view; G, palpal tibia, dorsal view. (Scales. A – D: 1 mm; E – G: 0.25 mm)

FIGURE 4. M. alticephalon sp. nov., male. A, body, dorsal view; B, chelicera and carapace, lateral view; C, chelicera and fang, ventral view; D, maxillae, labium, sternum and coxae; E, palp, ventral view; F, palp, lateral view; G, palp, dorsal view. (Scales. A – D: 1 mm; E – G: 0.25 mm)

FIGURE 20. M. hanoii, male. A, body, dorsal view; B, carapace, lateral view; C, maxillae, labium, sternum and coxae; D, chelicera and fang, ventral view; E, palp, ventral view; F, palp, lateral view; G, palpal tibia, dorsal view. (Scales. A, C – D: 0.50 mm; B: 1 mm; E – G: 0.25 mm)

FIGURE 10. M. borneensis, male. A, chelicera and carapace, dorsal view; B, abdomen, dorsal view; C, chelicera and carapace, lateral view; D, maxillae, labium, sternum and coxae; E, chelicera and fang, ventral view; F, cymbium, embolus and tegulum; G, palpal tibia, ventral view. (Scales. A – E: 1 mm; F – G: 0.25 mm)

FIGURE 30. M. maxillasa, male. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D, chelicera and fang, ventral view; E, palp, ventral view; F, palp, lateral view; G, palpal tibia, dorsal view. (Scales. A – D: 1 mm; E – G: 0.25 mm)

FIGURE 17. M. endoi sp. nov., male. A, body, dorsal view; B, chelicera and carapace, lateral view; C, chelicera and fang, ventral view; D, maxillae, labium, sternum and coxae; E, palp, ventral view; F, palp, lateral view; G, palpal tibia, dorsal view. (Scales. A – D: 1 mm; E – G: 0.25 mm)

FIGURE 28. M. markaha, male. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D, chelicera and fang, ventral view; E, palp, ventral view; F, palp, lateral view; G, palpal tibia, dorsal view. (Scales. A – D: 1 mm; E – G: 0.25 mm)

FIGURE 15. M. crassembolus sp. nov., male. A, body, dorsal view; B, maxillae, labium, sternum and coxae; C, body, lateral view; D, chelicera and fang, ventral view; E, palp, ventral view; F, palp, lateral view; G, palpal tibia, dorsal view. (Scales. A – D: 1 mm; E – G: 0.25 mm)

FIGURE 24. M. malayana, male. A, body, dorsal view; B, chelicera and carapace, lateral view; C, chelicera and fang, ventral view; D, maxillae, labium, sternum and coxae; E, palp, ventral view; F, palp, lateral view; G, palpal tibia, dorsal view. (Scales. A – D: 1 mm; E – G: 0.25 mm)

FIGURE 7. M. assimilis, female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D and F, internal structure of epigyne, ventral view; E, internal structure of epigyne, dorsal view. (Scales. A – C: 1 mm; D – E: 0.25 mm; F: 0.10 mm)

FIGURE 3. M. acromegalis sp. nov., female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D – E, internal structures of epigyne, ventral view; F, internal structure of epigyne, dorsal view. (Scales. A – C: 1 mm; D, F: 0.25 mm; E: 0.10 mm)

FIGURE 14. M. cornuta, female. A, body, dorsal view; B, maxillae, labium, sternum and coxae; C, body, lateral view; D and F, internal structure of epigyne, ventral view; E, internal structure of epigyne, dorsal view. (Scales. A – C: 1 mm; D – E: 0.25 mm; F: 0.10 mm)

FIGURE 33. M. melanocephala, female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D, E and G, internal structure of epigyne, ventral view (they are different specimens); F and H, internal structure of epigyne, dorsal view (F is the same specimen with E, and H with G). (Scales. A – C: 1 mm; D – H: 0.25 mm)

FIGURE 9. M. biseratensis, female. A, body, dorsal view; B, body, lateral view; C, maxillae, labium, sternum and coxae; D and F, internal structure of epigyne, ventral view; E, internal structure of epigyne, dorsal view. (Scales. A – C: 1 mm; D – E: 0.25 mm; F: 0.10 mm)

FIGURE 25. M. malayana, female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D – E, internal structure of epigyne, ventral view. (Scales. A – C: 1 mm; D: 0.25 mm; E: 0.10 mm)

FIGURE 21. M. hanoii, female. A, body, dorsal view; B, carapace, lateral view; C, maxillae, labium, sternum and coxae; D, internal structure of epigyne, ventral view. (Scales. A – C: 0.50 mm; D: 0.10 mm)

FIGURE 12. M. brevichelicera sp. nov., female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D and F, internal structure of epigyne, ventral view; E, internal structure of epigyne, dorsal view. (Scales. A – C: 1 mm; D – E: 0.25 mm; F: 0.10 mm)

FIGURE 23. M. lambirensis sp. nov., female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D – E, internal structure of epigyne, ventral view (D, E from the same specimen); F, internal structure of epigyne (different specimen from D, E), ventral view. (Scales. A – C: 1 mm; D, F: 0.25 mm; E: 0.10 mm)

FIGURE 29. M. markaha, female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D and F, internal structure of epigyne, ventral view; E, internal structure of epigyne, dorsal view. (Scales. A – C: 1 mm; D – E: 0.25 mm; F: 0.10 mm)

FIGURE 5. M. alticephalon sp. nov., female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D – E, internal structure of epigyne, ventral view. (Scales. A – C: 1 mm; D: 0.25 mm; E: 0.10 mm)

FIGURE 19. M. gedongensis, female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D – E, internal structure of epigyne, ventral view. (Scales. A – C: 1 mm; D: 0.25 mm; E: 0.10 mm)

FIGURE 31. M. maxillosa, female. A, body, dorsal view; B, chelicera and carapace, lateral view; C, maxillae, labium, sternum and coxae; D – E, internal structure of epigyne, ventral view. (Scales. A – C: 1 mm; D: 0.25 mm; E: 0.10 mm)