Phanocles ecuadoricus, Hennemann & Conle, 2024

Hennemann, Frank H. & Conle, Oskar V., 2024, Studies on Neotropical Phasmatodea XXVI: Taxonomic review of Cladomorformia tax. n., a lineage of Diapheromerinae stick insects, with the descriptions of seven new genera and 41 new species (Phasmatodea: Occidophasmata: Diapheromerinae), Zootaxa 5444 (1), pp. 1-454 : 147

12. Phanocles ecuadoricus sp. n. ( Figs. 51A , 55A , 58F , 59H , 61G–J , 66P–Q , 88H , 92G , 99D , 111A–B )

Remarks: For type data see description below.

Distribution: NW-Ecuador, Prvince Imbabura, Paramba [ZMUH]; NW-Ecuador, Province Pichincha, Puerto Quito [NHMUK]; NW-Ecuador, Province Pichincha, Union del Tuachi / Meija, Bastion del Diablo, 800 m [coll. FH]; NWEcuador, Province Pichincha, Cachabi [NHMW]; NW-Ecuador, Privince Pichincha, Mindo 1240 m [FH]; NW-Ecuador, Province Esmeraldas, Carondeled [ANSP]; NW-Ecuador, Province Esmeraldas, El Placer, 675 m [ANSP]; NW-Ecuador, Province Esmeraldas, Rio Verde [MNCN]; NW-Ecuador, Province Pichincha, Santo Domingo de los Colorados [MNHN]; NW-Ecuador, Province Pichincha; Mashpi Nature Reserve, 800-1000 m [OC].

Figures

FIGURE 51. Phanocles Stål, 1875 habitus (in scale): A. P. ecuadoricus sp. n. ♀ HT (Ecuador, Prov. Imbabura, Paramba), dorsolateral view [ZMUH]; B. P. falcatus sp. n. ♀ PT (Ecuador, Prov. Napo, Misahualli), dorsolateral view [coll. FH, No. 0919–1]; C. P. virgulatus (Redtenbacher, 1908) ♀ (Peru, Dept. Huanuco, Rio Lullapichis, Panguana) dorsolateral view [coll. FH, No. 0538–3]; D. P. brevipes sp. n. ♀ HT (Peru, Prov. Satip, Junín, Satipo), dorsal view [ANSP]; E. P. satyr (Redtenbacher, 1908) ♀ (S-Venezuela, San Esteban), dorsolateral view [ANSP].

FIGURE 55. Phanocles Stål, 1875: A. P. ecuadoricus sp. n. ♂ PT (Ecuador, Prov. Esmeraldas, Rio Verde), dorsolateral view [MNCN]; B. P. falcatus sp. n. ♂ PT (Ecuador, Prov. Napo, Misahualli), dorsolateral view [coll. FH, No. 0919–3]; C. P. maximus sp. n. ♀ PT (captive reared from Panamá, Prov. Coclé, Valle deAntón), dorsolateral view [coll. FH, No. 1360–3]; D. P. spectabilis sp. n. ♂ PT (captive reared from Costa Rica, Prov. Puntarenas, La Gamba), dorsolateral view [coll. FH, No. 0563–18]; E. P. ploiaria (Westwood, 1859) ♀ (Panamá, Prov. Coclé, Valle de Antón 650–850 m), dorsolateral view [coll. FH, No. 0705–12]; F. P. obtusus (Redtenbacher, 1908) ♂ (Venezuela, Aragua N.P., Rancho Grande), dorsolateral view [ANSP].

FIGURE 58. Phanocles Stål, 1875 head and prothorax: A. P. aequatorialis (Redtenbacher, 1908) ♀, lateral view [coll. FH]; B. P. costaricensis Hennemann, 2002 ♀, lateral view [coll. FH]; C. P. ploiaria (Westwood, 1859) ♀, lateral view [coll. FH]; D. P. maroniensis (Chopard, 1911) ♀ HT, lateral view [MNHN]; E. P. satyr (Redtenbacher, 1908) ♀ dorsolateral view [ANSP]; F. P. ecuadoricus sp. n. ♀ PT, lateral view [coll. FH]; G. P. maximus sp. n. ♀ PT, dorsolateral view [coll. FH]; H. P. spectabilis sp. n. ♀ PT, dorsolateral view [coll. FH]; J. P. spectabilis sp. n. ♀ PT, dorsolateral view [coll. FH]; K. P. subvolans (Redtenbacher, 1908) ♀, dorsolateral view [ANSP]; L. P. horni (Redtenbacher, 1908) ♀, dorsolateral view [coll. FH].

FIGURE 59. Phanocles Stål, 1875: A. P. mexicanus sp. n. ♀ HT (with small cephalic tubercles), lateral view of head and prothorax [MNHN]; B. P. mexicanus sp. n. ♀ PT (with prominent cephalic horns), lateral view of head and prothorax [MNHN]; C. P. mexicanus sp. n. ♀ HT, dorsal view of head, pro and mesonotum [MNHN]; D. P. pleurospinosus sp. n. ♀ HT, dorsolateral view of mesothorax with median segment showing the typical spines of the metapleurae [MNCR-A]; E. P. solidus sp. n. ♀ HT, lateral view of head and prothorax [ANSP]; F. P. maroniensis (Chopard, 1911) ♂, lateral view of head and prothorax [MNHN]; G. P. virgulatus (Redtenbacher, 1908) ♂, dorsolateral view of head and prothorax [coll. FH]; H. P. ecuadoricus sp. n. ♂ PT, dorsolateral view [MNCN]; J. P. falcatus sp. n. ♂ PT, dorsolateral view of head and prothorax [coll. FH].

FIGURE 61. Phanocles Stål, 1875 terminalia of ♀♀: A. P. burkartii (Saussure, 1868), lateral view [coll. FH]; B. P. burkartii (Saussure, 1868), dorsal view [coll.FH]; C. P.burkartii (Saussure, 1868), ventral view [coll. FH]; D. P.costaricensis Hennemann, 2002, lateral view [coll. FH]; E. P. costaricensis Hennemann, 2002, dorsal view [coll. FH]; F. P. costaricensis Hennemann, 2002, ventral view [coll. FH]; G. P. ecuadoricus sp. n. ♀ HT, lateral view [ZMUH]; H. P. ecuadoricus sp. n. ♀ HT, dorsal view [ZMUH]; J. P. ecuadoricus sp. n. ♀ HT, ventral view [ZMUH]; K. P. aequatorialis (Redtenbacher, 1908), lateral view [coll. FH]; L. P. aequatorialis (Redtenbacher, 1908), dorsal view [coll. FH]; M. P. aequatorialis (Redtenbacher, 1908), ventral view [coll. FH]; N. P. maroniensis (Chopard, 1911) ♀ HT, lateral view [coll. FH]; O. P. maroniensis (Chopard, 1911) ♀ HT, dorsal view [coll. FH]; P. P. maroniensis (Chopard, 1911) ♀ HT, ventral view [coll. FH]; Q. P. saussurei (Redtenbacher, 1908), lateral view [coll. FH]; R. P. saussurei (Redtenbacher, 1908), dorsal view [coll. FH]; S. P. saussurei (Redtenbacher, 1908), ventral view [coll. FH].

FIGURE 66.Phanocles Stål, 1875 terminaliaof ♂♂: A. P.maroniensis (Chopard, 1911), lateral view [MNHN]; B. P.maroniensis (Chopard, 1911), dorsal view [MNHN]; C. P. maroniensis (Chopard, 1911), ventral view [MNHN]; D. P. barbadosense sp. n. ♂ PT, lateral view [NHMW, © 2019 Natural History Museum Vienna, NOaS Image Collection / H. Bruckner]; E. P. barbadosense sp. n. ♂ PT, dorsal view [NHMW, © 2019 Natural History Museum Vienna, NOaS Image Collection / H. Bruckner]; F. P. barbadosense sp. n. ♂ PT, ventral view [NHMW, © 2019 Natural History Museum Vienna, NOaS Image Collection / H. Bruckner]; G. P. keratosqueleton (Olivier, 1792), ventral view (specimen from St. Vincent) [FSCA]; H. P. aequatorialis (Redtenbacher, 1908), lateral view [coll. FH]; J. P. acutecornutus sp. n. ♂ PT, lateral view [ZSMC]; K. P. acutecornutus sp. n. ♂ PT, dorsal view [ZSMC]; L. P. acutecornutus sp. n. ♂ PT, ventral view [ZSMC]; M. P. muticus (Brunner v. Wattenwyl, 1907), lateral view [coll. FH]; N. P. muticus (Brunner v. Wattenwyl, 1907), dorsal view [coll. FH]; O. P. muticus (Brunner v. Wattenwyl, 1907), ventral view [coll. FH]; P. P. ecuadoricus sp. n. ♂ PT, lateral view [MNHN]; Q. P. ecuadoricus sp. n. ♂ PT, dorsal view [MNHN].

FIGURE 88. Mesosternum: A. Phanocles burkartii (Saussure, 1868) ♀ [coll. FH]; B. Phanocles maximus sp. n. ♀ PT [coll. FH]; C. Phanocles maximus sp. n. ♂ PT [coll. FH]; D. Phanocles mexicanus sp. n. ♀ HT [MNHN]; E. Phanocles ploiaria (Westwood, 1859) ♀ [coll. FH]; F. Phanocles ploiaria (Westwood, 1859) ♂ [coll. FH]; G. Phanocles subvolans (Westwood, 1859) ♀ [ANSP]; H. Phanocles ecuadoricus sp. n. ♀ HT [ZMUH]; J. Phanocles maroniensis (Chopard, 1911) ♂ [MNHN]; K. Phanocles satyr (Redtenbacher, 1908) ♀ [ANSP]; L. Phanocles saussurei (Redtenbacher, 1908) ♀ [coll. FH]; M. Phanocles superbus sp. n. ♀ HT [MNHN]; N. Phanocles spectabilis sp. n. ♂ HT [coll. FH]; O. Phanocles falcatus sp. n. ♂ HT [ANSP]. Not in scale.

FIGURE 92. Praeopercular organ of ♀♀: A. Phanocles berezini sp. n. PT [coll. FH]; B. Phanocles horni (Redtenbacher, 1908) [coll. FH]; C. Phanocles chaperi (Redtenbacher, 1908) HT [MNHN]; D. Phanocles keratosqueleton (Olivier, 1792) St. Vincent [coll. ASPER, © Philippe Lelong]; E. Phanocles keratosqueleton (Olivier, 1792) Dominica [coll. ASPER, © Philippe Lelong]; F. Phanocles mutica (Brunner v. Wattenwyl, 1907) Trinidad [coll. ASPER, © Philippe Lelong]; G. Phanocles ecuadoricus sp. n. PT [ZMUH]; H. Phanocles maximus sp. n. [coll. FH]; J. Phanocles spectabilis sp. n. PT [coll. FH]. Not in scale.

FIGURE 99. Eggs (in scale): A. Phanocles aequatorialis (Redtenbacher, 1908) [coll. FH]; B. Phanocles costaricensis Hennemann, 2002; C. Phanocles cuzcoense sp. n. ex ovipositor HT [ANSP]; D. Phanocles ecuadoricus sp. n. PT [coll. FH]; E. Phanocles maximus sp. n. [coll. FH]; F. Phanocles mutica (Brunner v. Wattenwyl, 1907) [coll. FH]; G. Phanocles ploiaria (Westwood, 1859) Panamá, Prov. Coclé, Valle de Antón [coll. FH]; H. Phanocles rehni sp. n. ex ovipositor HT [ANSP]; J. Phanocles spectabilis sp. n. PT [coll. FH]; K. Phanocles vosseleri (Redtenbacher, 1908) [coll. FH]; L. Phanocles virgulatus (Redtenbacher, 1908) [coll. FH]; M. Phanocles keratosqueleton (Olivier, 1792) Martinique [coll. FH]; N. Phanocles keratosqueleton (Olivier, 1792) St. Vincent [coll. FH]; O. Phanocles chaperi (Redtenbacher, 1908) [coll. FH]; P. Phanocles falcatus sp. n. PT [coll. FH]. [Scale = 1 mm]

FIGURE 111. Live insects: A. Phanocles ecuadoricus sp. n. captive reared ♀ at Montréal Insectarium, Canada (© René Limoges); B. Phanocles ecuadoricus sp. n. captive reared ♂ at Montréal Insectarium, Canada (© René Limoges); C. Phanocles aequatorialis (Redtenbacher, 1908) captive reared ♀ from Durango, Ecuador.