Anatoma australis ( Hedley, 1903 )

Geiger, Daniel L. & Jansen, Patty, 2004, Revision of the Australian species of Anatomidae (Mollusca: Gastropoda: Vetigastropoda), Zootaxa 415 (415), pp. 1-35 : 9-14

Anatoma australis ( Hedley, 1903) : Figures 3–5 , 18

Scissurella australis Hedley, 1903: 329–330 , fig. 63.

Scissurella australis: Hedley, 1907: 288 .

Scissurella australis: Verco, 1910: 115 .

Scissurella australis: Thiele, 1912: 19 , pl. 3, fig. 1 [copy of figure in Hedley, 1903].

Schizotrochus australis: Cotton & Godfrey, 1933: 22–23 , pl. 1, fig. 10.

Schizotrochus australis: Iredale & McMichael, 1962: 30 .

Schizotrochus australis: Macpherson & Gabriel, 1962: 34 .

Scissurella (Anatoma) australis: Wilson, 1993: 46 .

Anatoma australis: Jansen, 1995 , 11.

Anatoma australis: Hickman, 1998 : fig. 15.52 A–B.

Anatoma australis: Jansen, 1999: 48 , figs. 4–6.

Anatoma “SWA”: Jansen, 1999: 50, figs. 16–18.

Type material. HOLOTYPE ( AMS C.16223), 3 x 2.5 mm (W x H). PARATYPES: 3 ( AMS C.170412).

Type locality. [5–8 mls.] off Port Kembla , [NSW,] 63–75 fms. [= 115–137 m; 34.465°S, 151.075°E] (OD) .

Etymology. Most likely referring to the occurrence in the southern hemisphere.

Description. Shell globular, medium size to large (3.6 mm). Protoconch 3/4 whorls, with flocculant sculpture, varix present, varix not connected to embryonic cap. Teleoconch I with spiral cord in area of selenizone, often with additional spiral cords, but no centrifugal cords, more than 0.5 whorls. Teleoconch II with up to 2.5 whorls. Sculpture on shoulder consisting of 1–4 spirals at the start of teleoconch II, approximately 12 spirals at aperture of fully grown specimens (2.5 mm), spirals on early part of teleoconch II much weaker than axials, at aperture at most as strong as axials; approximately 70–75 axials on shoulder and base of body whorl in fully grown shell. Base with approximately a dozen spiral cords. Spirals on base as strong as axials; spirals running over axials forming weak nodule. Umbilicus sloping continuously from base, narrow, deep. Selenizone at periphery, keels distinct, growth lines in selenizone not coordinated with axials; margins of slit converging towards apertural margin of shell. Aperture rounded, somewhat flared. Operculum multispiral, nucleus central, corneus, thin. Animal unknown.

Differential diagnosis. Anatoma tobeyoides n. sp. lacks the protoconch varix, on teleoconch I of fewer than 0.5 whorls, lacks a spiral cord in area of selenizone, and has fine centrifugal lines, but no true spiral sculpture.

Distribution. WA (Cape Naturaliste: 33.8°S to 11.5°S), NSW (Port Kembla: 34.5°S) and QLD (Swain’s Reef: 22.5°S), 27–1,463 m depth shells only, 176 m live.

Specimen Records.

QLD. 201 m, off S end Fraser Island , 27.952S, 153.851E ( AMS C.400802, 3) . 128 m, off Cape Moreton , 27S, 153.583E ( AMS C.400806, 17) . 115 m, NE of Cape Moreton light, 26.925S, 153.558E ( AMS C.400807, 17; AMS C.400810, 3) . 183 m, NNE of Cape Moreton , 26.875S, 153.6E ( AMS C.400809, 20+) . 128 m, E of Caloundra , 26.8S, 153.583E ( AMS C.400805, 20+) . 200 m, off Maroochydore , 26.687S, 153.64E ( AMS C.400808, 9) . 128 m, ESE of Noosa Heads , 26.567S, 153.667E ( AMS C.400804, 18) . 210 m, off Fraser Island , 24.965S, 153.622E ( AMS C.400818, 6) . 150 m, NE of Lady Elliot Island , 24.062S, 152.823E ( AMS C.400813, 4) . 175 m, E of Lady Musgrave Island , 23.853S, 152.575E ( AMS C.400814, 20+) . 132 m, Capricorn Channel , E of Lady Musgrave Island, 23.835S, 152.535E ( AMS C.400811, 21) . 284 m, Capricorn Channel, E of North West Island , 23.253S, 152.402E ( AMS C.400816, 4) . 155 m, Capricorn Channel , 23.143S, 152.277E ( AMS C.400815, 12) . 115 m, Capricorn Channel , 16.8 ml NE of North Reef, 23.14S, 152.205E ( AMS C.400817, 20+) . 78 m, SE of Swain Reefs , 22.518S, 152.71E ( AMS C.400812, 3) . 187 m, SE of Swain Reefs , 22.438S, 153.285E ( AMS C.400803, 20+) .

NSW. 115 m, 5–8 mls off Port Kembla , 34.465S, 151.075E ( AMS C.16223, 1; AMS C.170412, 3) . 183 m, 16 mls E of Wollongong , 34.417S, 151.25E ( AMS C.17988, 3) . 161 m, off Port Kembla , 34.333S, 151.3E ( AMS C.400796, 1) . 205 m, S of Port Hacking , 34.16S, 151.438E ( AMS C.346346, 1) . 750 m, off Sydney , 34.083S, 151.727E ( AMS C.402694, 11) . 79 m, off Sydney , 34.05S, 151.617E ( AMS C.400793, 3; C.402613, 1) . 150 m, 0ff Sydney , 33.967S, 151.483E ( AMS C.400792, 14) . 200 m, off Sydney , 33.872S, 151.667E ( AMS C.400798, 8) . 1463 m, 35 mls E of Sydney , 33.85S, 151.967E ( AMS C.26669, 4) . 109 m, off Port Jackson , 33.833S, 151.45E ( AMS C.400801, 1) . 176 m, off Sydney , 33.767S, 151.717E ( AMS C.400791, 20+, 2: complete) . 174 m, off Sydney , 33.717S, 151.767E ( AMS C.400794, 1) . 143 m, off Broken Bay , 33.717S, 151.667E ( AMS C.400795, 2; C.400797, 2) . 146 m, 22 mls E of Narrabeen, N of Sydney , 33.7S, 151.717E ( AMS C.25803, 11) . 132 m, off Norah Head , 33.333S, 151.933E ( AMS C.400799, 1) . 183 m, NE of Port Macquarie , 31.383S, 153.2E ( AMS C.37724, 2) . 185 m, 24 km E of Ballina , 28.827S, 153.855E ( AMS C.400800, 9) .

TAS. 113 m, N of Great Oyster Bay , 41.758S, 148.517E ( LACM 73–156 , 2 ) .

WA. 27 m, Sahul Banks , Timor Sea, 11.5S, 125.5E ( AMS 377600 , 1 ) . Point Quobba , N of Carnarvon, 24.483S, 113.417E ( AMS 379976 , 1 ) . 108 m, off North West Cape, 22.705S, 113.54E ( AMS 402644 , 1 ; AMS 402645 , 1 ) . 238 m, North West Shelf , ca 230ml W Roebuck Bay, 18.5S, 118.05E ( AMS 402676 , 1 ) . 238 m, SW of Cape Naturaliste, 33.742S, 114.435E ( AMS 402717 , 8 ) . 256 m, NW of Cervantes , 30.5S, 114.633E ( AMS 402718 , 10 ) . 155 m, NW of Bunbury , 33.25S, 114.617E ( AMS 402719 , 1 ) . 274 m, NW of Beagle Is , 29.717S, 114.283E ( AMS 402720 , 7 ) . 197 m, NW of Green Head , 29.967S, 114.45E ( AMS 402721 , 1 ) . 176 m, W of Garden Is , 32.262S, 115.112E ( AMS 402741 , 1 ) . 146 m, W of Dongara , 29.35S, 114.117E ( AMS 402742 , 1 ) . 183 m, W of Dongara , 29.142S, 113.913E ( AMS 402743 , 1 ) . 219 m, W.of Dongara , 29.183S, 113.9E ( AMS 402744 , 1 ) . 223 m, off Jurien Bay , 30.133S, 114.5E ( AMS 402745 , 1 ) . 183 m, NW of Beagle Is , 29.725S, 114.333E ( AMS 402746 , 1 ) . 200 m, NW of Bunbury , 33S, 114.617E ( AMS 402747 , 6 ) . 237 m, W of Green Head , 30.75S, 114.767E ( AMS 402748 , 1 ) . 160 m, off Rottnest Is , 31.65S, 115.08E ( AMS 402749 , 1 ) . 732 m, W of Rottnest Is , 31.083S, 114.767E ( AMS 402751 , 1 ) . 210 m, W of Garden Is , 32.25S, 115.117E ( AMS 402750 , 2 ) . 238 m, off Cervantes Is, 30.533S, 114.683E ( AMS 402752 , 2 ) .

Literature record. Off Wilson’s Promotory ( Macpherson & Gabriel, 1962).

Remarks. Jansen (1999) separated the Western Australian specimens of A. australis under the informal name A. SWA. SEM did not reveal any consistent differences between east and west coast specimens, hence, we reunite the Western Australian specimens under A. australis .

Figures

FIGURE 3. Anatoma australis. AMS C.400817. Capricorn Channel, 16.8 miles NE of North Reef, QLD, Australia. Scale bar shell = 200 µm. Scale bars spire and protoconch = 50 µm.

FIGURE 4. Anatoma australis. A. AMS C.400817. Capricorn Channel, 16.8 miles NE of North Reef, QLD, Australia. Second specimen from lot, see Figure 3. B. AMS C.400805. E of Caloundra, QLD, Australia. C. AMS C.400811. Capricorn Channel, E of Lady Musgrave Island, QLD, Australia. Scale bars shell = 200 µm. Scale bars spire and protoconch = 50 µm.

FIGURE 5. Anatoma australis. A. AMS C.402717. 238 m, SW of Cape Naturaliste, WA, Australia. B. AMS C.402720. 274 m, NW of Beagle Island, WA, Australia. C. AMS C.402721. 197 m, NW of Green Head, WA, Australia. Scale bars shell = 200 µm. Scale bars spire and protoconch = 50 µm.

FIGURE 18. Distribution maps of Australian species of Anatomidae. Circle with star: type locality. Solid dots: specimen records. Squares: literature records.