Coniceromyia valdesi, Kung, Giar-Ann, 2009

Kung, Giar-Ann, 2009, Four distinctive new Neotropical species of Coniceromyia Borgmeier (Diptera: Phoridae) with patterned-wings, Zootaxa 2273, pp. 49-58 : 54-57

Coniceromyia valdesi new species

( Figs. 4 , 11–12 , 16, 19–22, 26; Morphbank images 477868, 477869, 477870)

Recognition. C. valdesi is most easily differentiated from other patterned-wing Coniceromyia by the break or weakening in wing vein M2. The oval arrangement of ventral setae and lack of darkened pigment on the wing further distinguish this species from others.

FIGURES 15–16. Photographs, wings, dorsal view. 15. Coniceromyia sakaii new species. 16. C. valdesi new species. (Abbreviations: br=break or weakening of M2; ov=oval patch of setae.)

C. valdesi keys near couplet 12 in the Kung & Brown key (2000a). The wing patterning of C. valdesi is not due to darkening of the membrane, but rather to short, dense setae arranged in an oval, and the break or weakening in vein M2. C. valdesi lacks three striae on the wing, differentiating it from C. striativena Borgmeier. The lack of darkening on the wing membrane between the anterior margin and M1 prevent the second lead in the couplet from being followed.

Description. Body length 1.88–2.16 mm, mean 2.05 mm. Frons dark brown. Frontal height 1.05–1.13 frontal width, mean frontal ratio 1.08. Flagellomere 1 brown, pyriform. Palpus light brown. Scutum, scutellum, and pleuron dark brown to brown. Anepisternal setae absent. Legs mostly brown, femora darker than tibiae and tarsi. Foretibia with 2 dorsal to anterodorsal setae. Foretarsomere 1 broadened apically, with anterior excavation, covered in short, dense, orange setae; with long, anterobasal seta, seta approximately three-quarters length of foretarsomere 1; with narrow anteroapical process ( Fig. 4 ). Foretarsus tapered apically. Tarsomeres 2–4 of foreleg about as long as wide. In one specimen, tarsal ratio of foretarsus 2.17:1.00:1.00:1.00:1.00. Basal one-third of posterior face of hind femur with triangular, ventral group of FIGURES 17–22. Scanning electron micrographs, wings, low magnification and high magnification. 17–18. Coniceromyia sakaii new species, ventral view. 19–20. C. valdesi new species, dorsal view. 21–22. C. valdesi new species, ventral view. (Abbreviations: br=break or weakening of M2; f=fold in wing membrane; ov=oval patch of setae; vs=ventral setae.)

small, blunt peglike setae; arrangement of peglike setae tapered towards ventral margin ( Figs. 11–12 ). Hind tibia with one dorsal seta in basal half. Costa slightly thickened (Fig. 16). Wing vein R2+3 entirely absent. Wing membrane with fold visible across vein M2; posterior to M2 basally, anterior to M2 apically (Figs. 19–20). Venter of wing membrane with elongate oval arrangement of short, dense setae posterior to M2 (Figs. 16, 21– 22). Wing veins M1 and M2 parallel basally, diverging apically, with M1 anteriorly directed at apex; M2 relatively straight. Apical two-fifths of vein M2 narrowed; M2 with small break or weakening at narrowing (Figs. 16, 21–22). CuA1 relatively straight, curved anteriorly at apex. Costa length 0.42–0.46 wing length, mean 0.44. Halter white. Abdominal tergites brown to dark brown. Venter of abdomen brown or gray. Left surstylus of genitalia broad, truncate ( Fig. 26 ). Left hypandrial lobe dorsoapically directed, with large dorsal expansion.

Distribution. Colombia.

Derivation of species name. This species is named for my friend and colleague, Ángel Valdés.

Holotype. 3. COLOMBIA: Putumayo, PNN La Paya, Cab. Viviano, 0.12°S, 74.93°W, 30.xi–15.xii. 2001, 320 m, E. Lozano, Malaise trap, CAP-2794 [ LACM ENT 216018] ( IAVH).

Paratypes. COLOMBIA: Putumayo, PNN La Paya, Cab. Viviano, 0.12°S, 74.93°W, 43, 30.xi– 15.xii. 2001, 320 m, E. Lozano, Malaise trap, CAP-2794 ( IAVH, LACM).

Figures

FIGURES 1 – 4. Foretibiae and foretarsi, dorsal view. 1. Coniceromyia browni new species (foretibia in posterior view). 2. C. hoggi new species. 3. C. sakaii new species. 4. C. valdesi new species.

FIGURES 9 – 12. Scanning electron micrographs, hind femora, posterior view, low magnification and high magnification. 9 – 10. Coniceromyia sakaii new species. 11 – 12. C. valdesi new species.

FIGURES 23 – 26. Genitalia, left lateral view. 23. Coniceromyia browni new species. 24. C. hoggi new species. 25. C. sakaii new species. 26. C. valdesi new species.