Goniagnathus

Viraktamath, C. A. & Gnaneswaran, R., 2009, Three new species of Goniagnathus (Hemiptera: Cicadellidae) from the Indian subcontinent with description of a new subgenus, Zootaxa 2224 (1), pp. 51-59 : 53-54

Key to species of Goniagnathus from the Indian subcontinent (males)

1 Male pygophore without dorsal appendage, with or without caudal appendage.......................................................... 2

- Male pygophore with dorsal appendage ....................................................................................................................... 3

2 Aedeagus with pair of ventral processes exceeding shaft (Palaearctic, Oriental) .. G. (E.) guttulinervis (Kirschbaum)

- Aedeagus without pair of ventral processes ( India: Rajasthan) .......................... G. (T.) syncerus Dash & Viraktamath

3 Aedeagal shaft rather tubular, long and of uniform width............................................................................................ 4

- Aedeagal shaft constricted before apex; gonopore surrounded by sclerotized rim ending in a lateral short projection ( India: Gujarat, Maharashtra, Karnataka, Tamil Nadu) ................................... G. (T.) symphysis Dash & Viraktamath

4 Pygophore lobe strongly narrowed caudally with appendage bordering caudo-dorsal margin only............................ 5

- Pygophore lobe broad or broadened caudally, with appendage bordering both caudo-dorsal and caudal margins .. 11

5 Aedeagus with pair of processes ( Figs 9 , 16 ) .............................................................................................................. 6

- Aedeagus with two pairs of processes ( Figs 13 , 22, 23 ).............................................................................................. 7

6 Aedeagus with long processes at midlength ( Figs 16, 17 ); fused subgenital plates with truncate caudal margin ( Fig. 15 ) ( Nepal) ............................................................................................................................. G. (T.) nepalicus sp. nov.

- Aedeagal shaft with short curved processes at apex ( Figs 9, 11 ); fused subgenital plate convexly rounded ( Fig. 8 ) ( India: Karnataka) .................................................................................................................. G. (T.) anufrievi sp. nov.

7 Aedeagal shaft with pair of apical and asymmetrical median pair of processes ( India Gujarat, Maharashtra, Karnataka, Mizoram, Meghalaya, Kerala. Sri Lanka) ..................................................................... G. (T.) punctifer (Walker)

- Aedeagal shaft with two pairs of processes either near apex or midlength ..................................................................8

8 Aedeagal shaft with two pairs of processes near apex................................................................................................. 9

- Aedeagal shaft with two pairs of processes near midlength...................................................................................... 10

9 Aedeagal shaft with lateral processes slender and shorter than dorsal processes ( India: West Bengal, Meghalaya, Maharashtra, Karnataka, Kerala, Lakshadweep Islands) ......................................................... G. (T.) appellans Baker

- Aedeagal shaft with lateral processes stouter and longer than dorsal processes ( India: Orissa, Karnataka).................. ................................................................................................................. G. (T.) quadripinnatus Dash & Viraktamath

10 Aedeagal shaft with ventral pair of processes directed caudally, about as long as apical length of shaft beyond origin of processes ( Fig. 22 ); subgenital plate with truncate caudal margin ( Fig. 20 ) ( Sri Lanka) .......................................... .............................................................................................................................................. G. (T.) zeylanicus sp. nov.

- Aedeagal shaft with ventral pair of processes directed anteriorly, shorter than apical length of shaft beyond origin of processes ( Fig 13 ); subgenital plate with concave caudal margin ( India: Gujarat, Karnataka) ..................................... ........................................................................................................................... G. (T.) concavus Dash & Viraktamath

11 Fused subgenital plate with truncate caudal margin, style extending beyond plate; aedeagal shaft with pair of forked apical processes ( India: Gujarat, Maharashtra, Karnataka, Kerala, Tamil Nadu, Andhra Pradesh) .............................. ................................................................................................................................................... G. (T.) fumosus Distant

- Fused subgenital plate with caudal margin slightly convex, medially notched; styles not exceeding length of plate; aedeagal shaft with a pair of dorsal and an unpaired prong-like process ( India: Andhra Pradesh, Maharashtra, Karnataka, Pondicherry, Kerala. Sri Lanka) .................................................................................. G. (T.) nervosus Melichar

Figures

FIGURES 7–13. Species of Goniagnathus (Tropicognathus): G. (T.) anufrievi sp. nov. 7. Pygophore lobe, lateral view; 8. Fused subgenital plate and style, dorsal view; 9. Aedeagus and part of style, lateral view; 10, 11. Dorsal and ventral views of apex of aedeagus; 12. Female seventh sternite. G. (T.) concavus: 13. Aedeagus lateral view (both dorsal and ventral processes of aedeagal shaft are paired).

FIGURES 14–18. Goniagnathus (Tropicognathus) nepalicus sp. nov. 14. Pygophore lobe, lateral view; 15. Fused subgenital plate and style, dorsal view; 16. Aedeagus, lateral view; 17. Aedeagus, caudal view; 18. Female seventh sternite.

FIGURES 19–24. Goniagnathus (Tropicognathus) zeylanicus sp. nov. 19. Pygophore lobe, lateral view; 20. Fused subgenital plate and style, dorsal view; 21. Style, lateral view; 22. Aedeagus, lateral view; 23. Aedeagus, caudal view; 24. Female seventh sternite.