Mycomyiella irwini Väisänen

Gammelmo, Øivind, 2004, A review of the Afrotropical genus Mycomyiella Matile, 1973 (Diptera, Mycetophilidae, Mycomyini), with the description of six new species, Zootaxa 625, pp. 1-30 : 22-23

Mycomyiella irwini Väisänen

(Figs 15–18)

Mycomyiella irwini Väisänen, 1994: 21 , figs 38–42. Type locality: SOUTH AFRICA, Natal, Pietermaritzburg, Belfort; ZMH; male.

Material examined: SOUTH AFRICA, Natal, 75 km WSW Estcourt, Cathedral Peaks, 1700 m a.s.l., 26–31.xii.1979, 1 male, 1 female (paratypes), S. & J. Peck ( ZMH).

Diagnostic characters: The male groups with M. kaputuensis new species in having a forked gonostylus and in having the lobe of the gonocoxite reduced. It can be separated from M. kaputuensis in having a short tergite 9 without lobes.

Description

Male (n=1). Total length 3.18 mm.

Coloration: Head, palpomeres, and other mouthparts brown. Ocellar area dark brown. Face yellow, posterior parts of head brown. Antennae yellow­brown; scape, pedicel and first flagellomere yellow. Antepronotum yellow. Scutum brown to dark brown, with 3 yellow longitudinal bands. Anepisternum and preepisternum yellow. Scutellum light brown to brown. Laterotergite light brown to brown, mediotergite brown. Coxae yellow, femura, tibiae and tarsi light brown. Tergite 1 pale yellow, tergite 2 yellow, tergite 3 brown, tergite 4 brown with 2 large yellow lateral patches, tergites 5–6 brown, tergite 7 yellow. Sternites 1–4 yellow, sternites 5–6 brown, sternites 7 yellow. Terminalia light brown.

Head: Head 0.46 mm long; 0.47 mm wide. First flagellomere about 1.5 times as long as wide; second flagellomere as long as wide. Clypeus 0.14 mm long, with lateral setae. Ocellar area 0.14 mm wide, placed above upper margin of compound eye. Some setae in front of ocellar area. Posterolateral portion of frons densely setose, setae reaching below ocellar area. Face bare. Mouthparts moderately long. Palpomere 4 with few setae subbasally, palpomere 3 with several setae in basal two­thirds. Palpomere ratios (2–5) as: 1.00: 3.07: 2.89: 4.98.

Thorax: Total length 0.81 mm. Scutum 0.63 mm long, scutellum 0.18 mm long.

Legs: Leg measurements and ratios as in Tables 4 & 5.

Wing: Wing 3.18 mm long, 1.10 mm wide. Sc1 missing. Sc ending in R1 slightly distal to middle of small cell. Sc with 8 macrotrichia apically. R4/Rs­cell 1.0–2.0 times as long as wide. CuA fork slightly distal to M fork. Wing measurements are given in table 6.

Abdomen: Abdomen 2.54 mm long. Tergite 5 2.67 mm wide.

Male terminalia ( Figs 11 –18). Tergite 8 wide, with several long setae posterolaterally. Tergite 9 well developed, with long, bare lobes. Tergite 9 setose anterolaterally, bare medially. Gonocoxite setose laterally. Gonocoxite lobes reduced, with few setae. Gonocoxite lateral prolongation well developed and very prolonged (longer than lobes of tergite 9). Gonostylus slightly forked, with 4–5 setae. Aedeagus large, with long seta on posterior part, small setae present and evenly distributed. Sternite 8 pointed posteriorly.

Female (n=1). Total length 3.24 mm. As for males except:

Coloration: Tergite 4 brown to brownish. Tergite 7 yellow­brown with light brown lateral patches. Sternite 4 light brown. Sternite 7 yellow­brown with light brown lateral patch.

Head: Palpomere ratios (2–5) as: 1.00: 2.88: 2.76: 5.11.

Thorax: Total length 0.93 mm. Scutum 0.76 mm long, scutellum 0.16 mm long.

Wings: Wing 3.63 mm long, 1.13 mm wide.

Abdomen: Length of abdomen 2.67 mm. Tergite 5 2.49 mm wide.

Female terminalia: Cercus 2 elongated and pointed. Tergite 9 pointed on posterolateral corners. Gonocoxite 9 well developed. Sternite 10 normal or somewhat reduced.

Distribution: Known from Natal, South Africa only.

Figures

FIGURES 11 – 14. Mycomyiella ghanaensis new species. Male terminalia: 11 ventral; 12 dorsal; 13 sternite 8; 14 gonostylus. FIGURES 15 – 18. Mycomyiella irwini Väisänen, 1994. Male terminalia: 15 ventral; 16 dorsal; 17 sternite 8; 18 gonostylus.