Geodromicus convexicollis, Redtenbacher

Shavrin, Alexey V., 2022, The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae), Zootaxa 5213 (5), pp. 451-496 : 465

Redefinition of the convexicollis group

Diagnosis. Body medium-sized (4.25–7.90 mm); pronotum convex, moderately narrow or distinctly transverse, from widest anterior third gradually or sharply narrowed toward subparallel-sided short or elongate laterobasal portion; elytra slightly longer than broad ( G. (s.str.) malcolmi ) or distinctly broader than long, slightly or significantly broadened posteriad; aedeagus narrow or wide, with parameres slightly or significantly broadened anteriad, with weakly sclerotized internal sac and very narrow short or long flagellum.

Species included. G. (s.str.) convexicollis Luze, 1903 , G. (s.str.) kashmirensis Cameron, 1930 , G. (s.str.) lebedevianus Roubal, 1929 , G. (s.str.) malcolmi sp.n., G. (s.str.) macrothorax Kashcheev, 1999 , G. (s.str.) medvedevi Shavrin, 2009 .

Distribution. The species of the bodemeyeri group are distributed in the Palaearctic Region, and known from the Central Asia and the Himalayan Region.

Bionomics. The species of the convexicollis group inhabit banks of rivers and streams, and can be found in gravel, under stones, mosses, drift, etc. Specimens were collected at elevations from 700 to 4200 m a.s.l. from April to October.

Remarks. Previously, Bordoni (1984) included Geodromicus hauserianus (synonym of G. (s.str.) convexicollis ) to the rousi group ( Shavrin 2021). Based on the general shape of the apical portions of the parameres and weakly sclerotized internal sac with narrow flagellum, the convexicollis group is similar to the bodemeyeri group, from which it can be distinguished by the absence of paired sclerotized structures in apical and middle portions of the median lobe.

Key to convexicollis group

1 Elytra slightly longer than broad, slightly more than twice as long as pronotum. Median lobe narrow, with very wide parameres ( Fig. 64 ). Body length: 4.90 mm. Habitus as in Fig. 60 . India......................................... G. malcolmi

- Elytra as broad as or broader than long, distinctly less than twice as long as pronotum. Median lobe narrow or wide, with moderately narrow or wide parameres.................................................................... 2

2 Pronotum slightly broader (1.1/1.2) than long.............................................................. 3

- Pronotum distinctly transverse, 1.4/1.8 as broad as long...................................................... 4

3 Pronotum sharply narrowed toward long laterobasal margins. Parameres strongly broadened apically, distinctly longer than median lobe ( Fig. 52 ). Body yellow-brown to black, usually with wide and long pale macula on each elytron. Body length: 4.72–6.05 mm. Habitus as in Fig. 44–45 . Kyrgyzstan............................................ G. lebedevianus

- Pronotum from gradually narrowed toward moderately short laterobasal margins. Parameres slightly broadened apically, slightly exceeding apex of median lobe ( Shavrin 2009: fig. 1). Body brown, without maculae. Body length: 5.90–7.20 mm. Habitus as in Fig. 61 . Pakistan................................................................ G. medvedevi

4 Pronotum very convex, without or with indistinct longitudinal depression. Median lobe very narrow; parameres significantly exceeding apex of median lobe, with four long apical and three to four subapical setae ( Figs. 54, 56 ). Body length: 4.85–7.90 mm. Body yellow-brown to black. Habitus as in Fig. 57–59 . Kazakhstan, Kyrgyzstan................... G. macrothorax

- Pronotum less convex, without longitudinal depression. Median lobe wide; parameres slightly exceeding apex of median lobe, with four moderately short apical setae.................................................................... 5

5 Median lobe very wide, from apical third strongly narrowed apicad; apical portions of parameres moderately narrow ( Figs. 31, 33–36 ). Body length: 4.30–7.20 mm. Body brown to black. Habitus as in Fig. 38–39 . Middle Asia........ G. convexicollis

- Median lobe moderately narrow, from basal portion gradually narrowed apicad; apical portions of parameres distinctly broadened ( Figs. 46–49 ). Body length: 4.25–7.20 mm. Body reddish-brown to dark-brown Habitus as in Fig. 40–43 . Afghanistan, Pakistan, India................................................................................... G. kashmirensis

Figures

FIGURES 64–69. Aedeagus of Geodromicus malcolmi (Figs. 64–65: holotype) and G. affinis (Fig. 66: holotype; Figs. 67–68: India, Sonamarg; Fig. 69: Pakistan, Utrar): 64, 66–67, 69—parameral view, 65, 68—lateral view. Scale bars: 0.1 mm.

FIGURES 57–62. Habitus of Geodromicus: 57–59—G. macrothorax (Fig. 57: Kazakhstan, Chelkydysu; Fig. 58: Kazakhstan, Kokjota Mt.; Fig. 59: Kyrgyzstan, Grigoryevka), 60—G. malcolmi (holotype), 61—G. medvedevi (holotype), 62—G. affinis (holotype). Scale bars: 1.0 mm.

FIGURES 52–56. Aedeagus of Geodromicus lebedevianus (Figs. 52–53: Kazakhstan, Chichkan Valley) and G. macrothorax (Figs. 54–55: Kazakhstan, Chelkydysu; Fig. 56: Kyrgyzstan, Grigoryevka): 52, 54, 56—parameral view, 53, 55—lateral view. Scale bars: 0.1 mm.

FIGURES 40–45. Habitus of Geodromicus: 40—G. kashmirensis (lectotype), 41—G. similis (lectoype), 42—G. consors (holotype), 43—G. karakorus (holotype), 44–45—G. lebedevianus (Fig. 44: Kyrgyzstan, Chichkan Vallley; Fig. 45: Kyrgyzstan, Kara-Balta). Scale bars: 1.0 mm.

FIGURES 31–36. Aedeagus of Geodromicus convexicollis (Figs. 31–32: Kazakhstan, Kyrgyzsay Pass; Fig. 33: Kazakhstan, Kaskasu; Fig. 34: Kazakhstan, Ketmen Mnt.; Fig. 35: Tajikistan, Kulikulan; Fig. 36: Kyrgyzstan, Terskey-Alatoo Mts.): 31, 33–36—parameral view, 32—lateral view. Scale bars: 0.1 mm.

FIGURE 38–39. Habitats of Geodromicus convexicollis in Uzbekistan, Chatkal Mts., Iertash River.

FIGURES 46–50. Aedeagus of Geodromicus kashmirensis (Fig. 46: Afghanistan, Pagman; Fig. 47: India, Gulmarg; Fig. 48: India, Gya-Ladak; Figs. 49–50: India, Tangmarg Pir Panjal): 46–49—parameral view, 50—lateral view. Scale bars: 0.1 mm.