Palaeopoecilostola eocenica , Kania, Iwona & Krzemiński, Wiesław, 2012

Kania, Iwona & Krzemiński, Wiesław, 2012, A new species of Palaeopoecilostola Meunier, 1899 (Diptera: Limoniidae) from the Eocene Baltic amber, Zootaxa 3495, pp. 42-56: 43-45

publication ID 10.5281/zenodo.213542

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scientific name

Palaeopoecilostola eocenica

sp. nov.

Palaeopoecilostola eocenica  sp. nov.

( Figs 1–8View FIGURES 1 – 4View FIGURES 5 – 8)

Diagnosis. Antennae 16 -segmented, covered by long but not numerous setae, the base of each flagellomeres with two very elongated setae and rarely appearing shorter setae, usually reaching more than half the length of the elongated setae; outer dististylus wide and short, strongly expanded at the end, with short and thick setae; inner dististylus wide at the base, tapered at the top.

Description. The body length: 12.5 mm.; wing length 6.9 mm, width 4.1 mm, antennae 2.6 mm long. Head ( Fig. 1View FIGURES 1 – 4): slender with characteristic huge eyes; antennae (Figs 1,2,5) shorter than mid-length of the abdomen, usually pale; shape almost cylindrical, small; pedicel laterally swollen, almost cylindrical in shape; antennal flagellomeres narrow, elongate, cylindrical, covered by tiny, wispy setae, each flagellomere with two elongated setae, additionally, a few 2–4 setae are visible on all flagellomeres, the same length or slightly longer than the length of the flagellomere on which they occur; the last segment almost as long as the sub-apical one, slightly shorter.

Palpi ( Fig. 6View FIGURES 5 – 8): 4 -segmented, the last segment subtly longer than the penultimate one.

Wings (Figs 1,4,7): strongly dark, stigma clearly separated; Sc long, ending a little bit past bifurcation of vein Rs into R 2 + 3 + 4 and R 5; Rs slightly longer than R 2 + 3 + 4; R 2 + 3 + 4 two times as long as R 3; R 1 ending opposite proximal ¼ of R 3 length; r-r (R 2) almost before the end of R 1; R 4 twice as long as R 3; discal cell small, rectangular; M 1 about 1 / 3 longer than petiola, cross vein m-cu behind the midpoint of discal cell base and approximately as long as d-cell base; A 1 straight and long; A 2 sinuous for half of its length, the distal part of this vein strongly curved towards the wing’s edge.

Hypopygium ( Figs 1, 3View FIGURES 1 – 4, 8View FIGURES 5 – 8): elongated, basistylus covered by wispy, elongate setae; outer dististylus wide, strongly expanded at the end, rather short with short and thick setae at outer edge; inner dististylus expanded at the base, narrow at the apex.

Age and occurence. Baltic amber, Upper Eocene. The Baltic amber is aged within the range of 38–47 Ma ( Ritzkowski 1997; Perkovsky et al. 2007). Absolute dating analyses of glauconites from Sambia Peninsula showed that the “blue earth” formation (amber bearing Prussian Formation) is allocated to the Middle Eocene (Lutetian: 44.1 ± 1.1 Ma) and is thus significantly older than previously assumed ( Wappler 2003, 2005). Also Weitschat & Wichard (2010) suggested older age of the Baltic amber. However, assumptions on the Middle Eocene (or older) age of Baltic amber was argued by Perkovsky et al. (2007), and the Upper Eocene (Bartonian/Priabonian: 37.7 ± 3 Ma) age of Prussian Formation is preferred by these authors.

Etymology. Specific epithet after the geological period—the Eocene. Material examined. Holotype, male. No. 19931, Coll. Museum of the Earth Polish Academy of Sciences, Warsaw, Poland.