Retoka curvipes, Kondorosy, 2008

Kondorosy, Előd, 2008, A revision of the Entisberus group (Hemiptera: Heteroptera: Rhyparochromidae), Acta Entomologica Musei Nationalis Pragae 48 (2), pp. 591-610 : 601-602

Retoka curvipes sp. nov.

( Figs. 9-10 , 19 )

Type locality. Malaysia, Pahang, Cameron Highlands, Tanah Rata.

Type material. HOLOTYPE: J, ‘ MALAYSIA West: PAHANG / Cameron Highlands / TANAH RATA, 1200-1500m / 3. II. – 19. II. 2005, / Cechovsky Petr lgt’ ( NHMW) . PARATYPE: ♀, same label data as holotype ( NHMW).

Description. Structure. Body relatively wide, shining, clavus and endocorium dull; sparsely covered with moderately long hairs and densely with additional very minute hairs; rather densely and strongly punctate.

Head moderately long (about 1.5 times as long as high), in lateral view at base of clypeus strongly declivent, densely punctate, punctures partly confluent (having the appearance of very narrow mesh-like carinae). Eyes large, bare, strongly protruding but not pedunculate, laterally surpassing anterolateral angles of pronotum, not projecting backwards. Hind margins of eyes situated close to pronotum but not touching it. Ocelli well developed, adjacent with pronotum. Antenniferous tubercles well developed, placed far from eyes, projecting laterad. Bucculae small, apically rounded, gradually narrowed posteriad, protruding from ventral outline of head in lateral aspect. Antennae slender, segments nearly cylindrical, with dense decumbent pubescence and rather sparse, long, prominent hairs; segment I surpassing apex of clypeus by less than one fourth of its length; segment II longer than segment I; remaining segments shorter than segment II (relative lengths of segments highly variable). Labium very short, segment I attaining anterior margin of eye, segment II longest, reaching base of pronotal collar, remaining segments shorter than segment I, labium slightly surpassing fore coxae.

Thorax. Pronotum bell-shaped, convex, anterior part with very dense and posterior lobe with sparser punctation. Pronotal surface uneven, posterior lobe with small impunctate tubercles, humeral angles strongly elevated, smooth, median groove shallow and wide. Pronotal collar indistinct, with two indistinct rows of punctures. Transverse impression well visible, situated at middle of pronotum. Lateral margins narrowly explanate, not laminate at level of transverse impression, abruptly terminating posteriorly and forming a tooth-like projection at middle part of humeral angles, reaching collar anteriorly. Basal margin of pronotum convex in front of scutellum, laterally depressed, anterior margin straight. Scutellum triangular with Y-shaped elevation, medially strongly elevated (this part about as high as pronotum, apex about one third of total length of scutellum), reaching neither basal nor apical angles, basal arms weak, directed posteriad towards basal angles, apical arm smooth. Hemelytra well developed, reaching apex of abdomen. Clavus with three regular rows of punctures. Corium with two rows of punctures along claval margin, lateral of them smooth to corial furrow, mesocorium with dense, irregularly distributed punctures, reflexed costal margin impunctate. Costal margin of corium convex at base, concave at level of scutellar apex, convex apically. Apical corial margin S-shaped, strongly concave on inner part. Membrane with four well visible veins. Sternum evenly punctate. Metasternal scent gland area small, occupying less than one third of metasternum. Legs slender, fore femur with two rather strong spines (male) or some fine teeth (female), fore tibia strongly curved around its midlength in male, posterior part with three teeth; straight and simple in female.

Abdominal venter without median keel. Sutures between sternites II-VI longitudinally striolate.

Male genitalia. Paramere of the usual ‘ Drymini type’ with inner lobe relatively wide ( Fig. 19 ). Parameres of the near relative R. gibba and R. esakii are shown on Figs. 17 and 18 .

Colouration. Body fuscous. Posterior pronotal lobe, clavus, base and inner margin of hemelytra, five patches on membrane, femora, base of tibiae and antennal segment I except apex dark brown. Antennal segment IV and tubercles on posterior pronotal lobe brown (the latter yellowish in the paratype). Two small spots on base and apex of clavus, three corial patches (a yellowish spot on endocorium at end of clavus, two transversal streaks on corium at one and two thirds of its total length), membrane, antennal segments II and III, labium, apices of tibiae and tarsi yellow.

Measurements (all in mm; J holotype first, ♀ paratype in parentheses). Total body length 3.10 (3.20). Head: length 0.50 (0.48), width 0.58 (0.60), interocular space 0.42 (0.45). Pronotum: length 0.85 (0.80), width at base (= total width) 1.30 (1.35). Scutellum length 0.52 (0.60), length of claval commissure 0.30 (0.28). Lengths of antennal segments I 0.30 (0.28), II 0.50 (0.45), III 0.38 (0.35), IV 0.35 (0.40). Lengths of labial segments I 0.28, II 0.32, III 0.28, IV 0.22.

Differential diagnosis. Retoka curvipes sp. nov. is very similar to R. callosa but differs from the latter in the following characters (character states of R. callosa are given in parentheses): fore femur of male with two rather strong spines, fore tibia strongly curved, armed with some spines on inner side near apex (male fore femur only with minute spines, fore tibia straight, unarmed); female with a few minute spines on fore femur (female fore femur unarmed).

Etymology. The species epithet is composed of the Latin adjectivum curvus (= curved, bent) and the Latin noun pes (= foot), referring to the very characteristic curved fore tibia of the male.

Distribution. Malaysia (Pahang).

Figures

Figs. 9-12. Habitus of Retoka species (9, 11-12 – body in dorsal view; 10 – detail of right anterior leg of male holotype). 9-10 – R. curvipes sp. nov.; 11 – R. stysi sp. nov.; 12 – R. minuta China, 1935.

Figs. 14-21. Parameres. 14 – Entisberus archetypus Distant, 1903 (specimen from Vietnam: Yen Bai Province); 15 – E. bergrothi sp. nov.; 16 – Retoka hirsuta sp. nov,; 17 – R. gibba (Zheng, 1981); 18 – R. esakii (Slater & Hidaka, 1958); 19 – R. curvipes sp. nov.; 20 – R. stysi sp. nov.; 21 – R. paraminuta sp. nov.