Paramacroxiphus bifasciatus, Ingrisch, 2008

Ingrisch, Sigfrid, 2008, Revision of the genera Paramacroxiphus C. Willemse 1961 and Pseudomacroxiphus C. Willemse 1961 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini), Zootaxa 1755 (1), pp. 1-34 : 11

Paramacroxiphus bifasciatus sp. n.

Figs. 6 , 12 , 32 , 40 , 46 , 52 , 95

Holotype (male): Indonesia: Papua, Neth. Ind.-Amer. New Guinea Exped., Bernhard Camp , 50m, 7.XI.1938, lg. J. Olthof, labelled: " Paramacroxiphus (?) bifasciatus sp. n. " . Holotype in Museum Zoologicum Bogoriense [Cibinong], Indonesia .

Description. Fastigium verticis: eye length = 1.3: 2.0 mm; projecting 0.7 mm in front of eyes. Femora with the following number of spines on ventral margins: profemur 7 external, 6–7 internal; mesofemur 7 external, 4 internal; postfemur 15–16 external, 13–14 internal.

Male. Stridulatory file slightly sinuate, circa 2.84 mm long; teeth spaced near base, narrowing towards apex ( Fig. 12 ); with about 141 teeth or 49.6 teeth per mm; in middle with 41.0 teeth per mm; in basal half with 33.5 teeth per mm. Tenth abdominal tergite slightly globose at base with a faint indication of a medial carina; apical area depressed; apex bilobate and roundly excised in between ( Fig. 32 ). Epiproct rounded-triangular, small, dorsal surface depressed. Cerci cylindrical but little dorso-ventrally compressed; internal margin carinate in basal area, thereafter widening to a large, rounded, dorsally concave projection until apex; apical margin curved and with a small tooth on ventral margin ( Fig. 32 ). Subgenital plate with lateral areas rather little sloping; with a medial and two rounded lateral carinae; lateral carinae prolonged into long, subcylindrical, apical projections, apex roundly excised in between; styli small inserted at the tip to the projections ( Fig. 40 ).

Titillators separate, with basal parts long-ovoid and slightly concave surface; apical parts sheathed ( Figs. 46 , 52 ). The semi-membranous sheaths circa ovoid in general outline, apico-internal surface dark with fine striation, dorsal rim pale crenulated, ventral area pale with fine striation gradually transiting into hyaline sacculi; sheaths of both sides connected in middle; on dorso-external side connected with a bilobate hyaline structure. Baso-lateral sclerites large, elongate, tri-carinate.

Female. unknown.

Coloration. Brownish testaceus. Frons with two blackish bands from antennal scrobae to and including mandibles; antennal scrobae, fastigium frontis, scapus and pedicellus partly darkened ( Fig. 6 ). Pronotum with a small dark spot at humeral sinus. Anterior tibia with a small dark spot below tympanum. Tegmen maculated with dark and pale cells; veins always pale; dorsal area pale. Antennae with spaced dark rings. Spines pale with dark tips.

Measurements of male: body 27; pronotum 9.8; tegmen 33; postfemur 22 mm.

Diagnosis. The new species is easily recognisable by the face with a pair of black vertical bands. This pattern it shares only with P. tessellatus . Of the latter only the female is known, of P. bifasciatus only the male. In general characters, both species differ by the shape of the frons, which is narrower in P. bifasciatus (1.62 times higher than wide) than in P. tessellatus (1.50 times higher than wide), with less convex lateral margins; the wings are comparatively shorter in P. bifasciatus (index tegmen: hind femur = 1.5 in P. bifasciatus , 2.0 in P. tessellatus ). The dark pattern on pronotum and hind femur is weak and less expanded in P. bifasciatus . From males of other species, the new species is easy to separate by the characteristic shapes of cerci, subgenital plate and titillators.

Etymology. The name of the new species refers to the vertical bands on the face.

Figures

FIGURES 1–6. Habitus (1–2), ovipositor (3), and face (4–6) of Paramacroxiphus species: 1–4, P. aberrans (1, 4, male paratype; 2–3, female paratype); 5, P. tessellatus female holotype; 6, P. bifasciatus male holotype.

FIGURES 11–15. Stridulatory files on underside of left tegmen of Paramacroxiphus males: 11, P.armatus holotype; 12, P. bifasciatus holotype; 13, P. brunneus holotype; 14, P. elongatus holotype; 15, P. irregularius holotype.

FIGURES 29–34. Male abdominal apex of Paramacroxiphus species in dorsal view (31, in dorso-lateral view; 34, left cercus): 29, P. brunneus holotype; 30, P.armatus holotype; 31, P. elongatus holotype; 32, P. bifasciatus holotype; 33–34, P. irregularius holotype. The arrow points at the baso-internal tooth of the cercus. 10t tenth abdominal tergite; ce cercus; pa process of paraproct; sg subgenital plate.

FIGURES 35–44. Male subgenital plate (35–43) and stridulatory area of right tegmen (44) of Paramacroxiphus species: 35, P. elongatus holotype; 36, P. irregularius holotype; 37, P.rufus holotype; 38, P. securiformis holotype; 39, P.armatus holotype; 40, P. bifasciatus holotype; 41, P. maculatus holotype; 42, P. aberrans holotype; 43, P. brunneus holotype; 44, P.aberransparatype. Abbreviations: m mirror, s scrapper, sp field of spinules.

FIGURES 45–49. Male titillators of Paramacroxiphus species in ventro-distal view: 45, P. brunneus holotype; 46, P. bifasciatus holotype; 47, P. irregularius holotype; 48, P. elongatus holotype; 49, P.armatus holotype (surface structure indicated at apex). Baso-lateral sclerite only shown on one side. Abbreviations: ap apical part of titillator; bp basal part of titillator; br bristles; bs baso-lateral sclerite; m membranous part of phallus.

FIGURES 50–56. Male titillators of Paramacroxiphus species in lateral view (50, 53), dorso-proximal view (51–52, 55– 56) and detail of apical part (54): 50–51, P.armatus holotype; 52, P. bifasciatus holotype; 53–54, P. brunneus holotype; 55, P. elongatus holotype; 56, P. irregularius holotype. Abbreviations: ap apical part of titillator; bp basal part of titillator; br bristles; bs baso-lateral sclerite; m membranous part of phallus. Scale bars = 1 mm.(ZMB); 1 female, do. 67, Pionierlager, 1.v.–30.vi.1912, Bürgers (ZMB); 1 female, do. [label unreadable], Bürgers (ZMB).