Soyedina vallicularia ( Wu, 1923 )

Grubbs, Scott A. & Baumann, Richard W., 2023, The Nemourinae (Insecta, Nemouridae) of the eastern Nearctic, Zootaxa 5306 (1), pp. 1-53 : 42

22. Soyedina vallicularia ( Wu, 1923)

Valley Forestfly

http://lsid.speciesfile.org/urn:lsid: Plecoptera .speciesfile.org:TaxonName:6084

( Figs. 145–150 )

Nemoura vallicularia Wu 1923:58 . Holotype male ( Cornell University Insect Collection ), Ithaca, (Tompkins Co.), New York, USA

Nemoura vallicularia Needham & Claassen, 1925:215

Nemoura (Soyedina) vallicularia Ricker, 1952:53

Soyedina vallicularia Illies, 1966:248

Soyedina vallicularia Zwick, 1973:357

Nemoura (Soyedina) vallicularia Hitchcock, 1974:110

Soyedina vallicularia Baumann, 1975:67

Soyedina vallicularia Stewart & Stark, 1988:188

Soyedina vallicularia Stewart & Stark, 2002:215

Soyedina vallicularia Grubbs, 2006:40

Soyedina vallicularia Stewart & Stark, 2002:215

Soyedina vallicularia Stark, 2017:215

Soyedina vallicularia Grubbs & Baumann, 2019:227

Distribution. Canada: LB, NS, ON, PQ. USA; CT, IA, IL, IN, KY, MA, MD, ME, MI, NY, OH, PA, TN, VA, WI, WV ( DeWalt et al. 2022)

Male. Macropterous. Body length 4.7–5.1 mm, forewing length 6.8–8.1 mm (n = 16). Gills absent. Cerci simple and unmodified. Paraprocts with two lobes, inner lobes lightly sclerotized and inconspicuous, outer lobes sclerotized, in basal half bulbous, tapered and straight-sided in apical half ( Fig. 149 ). Epiproct asymmetric, typical for genus ( Fig. 145 ); ventral sclerites partially recurved over abdomen, distinctly asymmetric, apically narrowly triangular ( Fig. 145 ); dorsal sclerites distinctly asymmetric, appearing scaly at higher magnifications ( Figs. 145‒ 148 ); open apically, inner member heavily sclerotized, flanged apically, lip asymmetric with apical half longer than basal portion ( Figs. 145‒146 ).

Female. Macropterous. Body length 5.7–8.2 mm, forewing length 7.8.– 9.7 mm (n = 20). Gills absent. Cerci simple and unmodified. Subgenital plate well-developed, subtriangular in shape; base broad, extending from posterior of seventh sternum entirely over eighth sternum and approximately ¼ over ninth sternum; terminating in broadly-rounded apex ( Fig. 150 ).

Larva. Described in full (see Stewart & Stark 2002 and all references therein).

Comments. Soyedina vallicularia is the most widespread amongst the eastern Nearctic Soyedina , including the Appalachian Mountains, Great Lakes region , and Ohio River and upper Mississippi River basins ( Grubbs & Baumann 2019).

Figures

FIGURES 145‒150. Soyedina vallicularia. 145, USA, Pennsylvania, stream into Traverse Creek, epiproct, ventral view; 146, USA, Maryland, tributary to Trading Run, epiproct apex, ventral view; 147, USA, New York, Kenney Brook, epiproct, lateral view; 148, USA, Maryland, tributary to Fifteenmile Creek, epiproct, caudal view; 149, USA, Kentucky, Spout Spring, outer paraproct lobe, lateral view. 150, USA, Pennsylvania, tributary Traverse Creek, female, abdominal terminalia, ventral view. ds = dorsal sclerite, im = inner member, opl = outer paraproct lobe, vs = ventral sclerite.