Ecnomina batyle Neboiss

Cartwright, David I., 2008, A review of the Australian species of Ecnomina Kimmins and Daternomina Neboiss (Trichoptera: Ecnomidae), Zootaxa 1774 (1), pp. 1-76 : 38

Ecnomina batyle Neboiss

Figs 75–77 , 174

Ecnomina batyle Neboiss, 1977: 59 , figs 272–276. – Neboiss 1986:153.

Diagnosis. Ecnomina batyle is a very distinctive species because it is the only species of Ecnomina which lacks inferior appendages but has a distinctive slender, bifid process (phallic guide) arising ventrally from near the base of the phallus.

Description. (Revised after Neboiss 1977). Head, body and wings brownish, abdomen paler ventrally; wings similar to E. legula ( Fig. 3 ). Forewing length about 3.0–3.2 times width: male 3.4–4.7 mm, female 4.0– 4.6 mm. Forewing fork 2 relatively long, sessile, length about 1.2–1.4 times length of fork 3; fork 3 relatively long, with short footstalk, length fork greater than 3.5 times length footstalk, footstalk length about 0.3–1.4 times length cross-vein m, r-m and m not contiguous at fork 3 by about 0.3–1.0 times length cross-vein m; fork 4 present, similar length to fork 3; fork 5 long, length about 1.8 times length of fork 4. Hindwing length about 3 times width, fork 2 sessile, length about 1.4–1.6 times length of fork 3.

Male. Tergum X membranous, sclerotised mesal process of tergum X short ( Fig. 77 ). Superior appendages in lateral view, sub rectangular, length about 3 times width ( Fig. 75 ); in dorsal view, length about 3 times width, expanded slightly meso-distally ( Fig. 77 ). Phallus generally tube-like, dilated and upturned distally; with a slender, bifid process (phallic guide) arising ventrally from near the base of the phallus. Inferior appendages absent ( Figs 75, 76 ).

Female. Genitalia with a single elongate and relatively slender mesal process on sternite VIII; segment IX relatively long, tapered slightly distally, segment X relatively short and segments IX and X relatively slender, with a few pairs of hairs on segment X ( Fig. 174 ).

Material examined: Tasmania. Holotype male, Wilmot River, 10 km S of Forth, 13 Dec 1974, A.N. (NMV, T- 4861), Paratypes, 12 males (specimen PT-475 figured), 12 females (specimen PT-476 figured), collected with holotype (NMV). Other material. Tasmania. 5 males, 25 females, Apsley R., 5 km NW of Bicheno, UV lt, 20 Dec 1988, J. Jackson. Other Tasmanian records refer to Neboiss (1977, 2003). Victoria. 5 males, 16 females, Yarra R., Warburton East, 17 Feb 1976, A.N.; 2 males, 20 females, Yarra R. nr McMahons Ck, 19 Feb 1976, A.N.; 1 male, Tanjil R. nr Old Tanjil, 38°3.2'S, 146°11'E, 5 Feb 1980, Latrobe C Survey; 1 male, Thomson R., Knappings Clearing, 9 Mar 1979, NMV Survey; 13 males, 16 females, same loc., 11 Feb 1980, Wells and McNamara; 4 males, 9 females, same loc., 9 Dec 1979, Wells and Bailey; 1 male, Thomson R., 7 km NNW Walhalla, Narrows Gauging Stn, 4 Mar 1980, NMV Survey; 18 males, 35 females, Bells Clearing, 6 km S of Aberfeldy, 9 Feb 1977, A.A. Calder; 1 male, same loc., 25 Feb 1978, NMV Survey Dept; 35 females, Howqua R., 12 km S of Merrijig, 18 Jan 1978, J.Blyth; 3 males, 1 female, King R., West Branch, 10 km SE of Tolmie, 2 Feb 1984, A.N.; 1 male, 7 females, Wonnangatta R., 37°09'S, 146°46'E, E of Mt Howitt, 11 Feb 1981, J.Blyth; 1 male, Porepunkah, 27, Jan 1957, A.N.; 4 males, 6 females, Cobungra R. at Anglers Rest, 15 Jan 1982, A.N.; 1 male, Mitta Mitta, 4 Feb 1961, E.M.; 3 males, 11 females, Mitta Mitta R., 8 km NE Benambra, 5 Feb 1974, A.N.; 1 male, Dart R., 11 Feb 1973, Dartmouth Survey; 11 males, 12 females, Gibbo R., Morass Ck jn, 1 Feb 1974, A.N.; 4 males, 6 females, Gibbo R. at Exhibition Ck jn, 20 km N of Benambra, 16 Jan 1982, A.W.; 2 females, Macalister-Caledonia R. jn, 16 Feb 1977, A.A. Calder; 1 male, Suggan Buggan, 24 Jan 1966, ND.; 1 male, Wellington R., 23 km NNE Licola on Tamboritra Rd, 25 Feb 1978, NMV Survey Dept; 2 males, 5 females, Wellington-Carey R. jn, 15 Feb 1977, A.A. Calder; 1 male, Noorinbee, 23 Nov 1965, A.N.; 1 male, Cann R., 33 km N of Cann River, 37°25'S, 149°11'E, 5 Feb 1987, K. Walker; 1 male, 3 females, Genoa R. at Wangarabell, 8 Feb 1980, A.W. New South Wales. 2 males, 2 females, Jindabyne, 21 Feb 1969, A.N.; 5 males, 6 females, Shoalhaven R. nr Braidwood, 2 Dec 1960, C.N. and A.S. Smithers (AM); 1 male, 4 females, same loc. and collectors, 26–30 Nov 1962 (AM); 1 male, 3 females, U. Kangaroo Valley, 24 Nov 1960, E.F. Riek (ANIC); 3 males, ‘Mooraback’, Werrikimbe NP, 980 m, 6 Dec 1986, G. Theischinger. Remarks. Ecnomina batyle is relatively common and widespread species throughout Tasmania, Victoria and eastern New South Wales (latitudinal range 31°15' - 43°38'S). Neboiss’s (1977) figures have been redrawn to allow direct comparisons and to accompany the description that is revised in light of new interpretations of Ecnomina genitalic and wing structures.

Figures

FIGURES 75–83. Ecnomina spp. Male genitalia in lateral, ventral and dorsal views. 75–77: Ecnomina batyle Neboiss. 78–80: Ecnomina vega Neboiss. 81–83: Ecnomina boogoo sp. nov. pro. IX, process on ninth abdominal segment. Figures 84–89. Ecnomina spp. Male genitalia in lateral, ventral and dorsal views. 84–86: Ecnomina attunga sp. nov. 87– 89: Ecnomina spinosa Kimmins.

FIGURES 168–174. Ecnomina spp. 168–170: Ecnomina sheldoni sp. nov. male genitalia in lateral, ventral and dorsal views. 171–172: Ecnomina merki sp. nov. 171: Head and thorax, dorsal view. 172: Wings, section of forewing and hindwing. 173–174: Female genitalia in ventral view. 173: Ecnomina scutica sp. nov. 174: Ecnomina batyle Neboiss.

FIGURES 1–6. 1–2. Daternomina irrorata (Kimmins). 1: Forewing and hindwing. 2: Head and thorax, dorsal view. 3– 6. Ecnomina spp. 3. Ecnomina legula Neboiss, forewing and hindwing. 4–5. Ecnomina scutica sp. nov. 4: Head and thorax, dorsal view. 5: Wings, section of forewing and hindwing. 6. Ecnomina spinosa Kimmins, head and thorax, dorsal view. n., nygma; o.w, occipital wart.