Oryctophileurus Kolbe, 1878,

Perger, Robert & Grossi, Paschoal Coelho, 2013, Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurin, ZooKeys 346, pp. 1-16: 2-4

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Oryctophileurus Kolbe, 1878


Oryctophileurus Kolbe, 1878 

Type species.

Phileurus nasicornis  Burmeister, 1847 (original combination) (Fig. 3).

Species of Oryctophileurus  are distinguished from other Phileurini  by a combination of the following apomorphic characters: a single cephalic horn with lateral carina, pronotal cavity with ocellate punctures and two teeth or tubercles close to the anterior pronotal margin, teeth vertically positioned at about the same level as the outer eye margins.

Taxonomy and discussion.

The following characters were cited by Endrödi (1985) for Oryctophileurus  , Amblyodus  , Microphileurus  Kolbe, Metaphileurus  Kolbe, Trioplus  Burmeister, and Goniophileurus  Kolbe and are here not considered as generic: outer side of mandible tridentate; antenna 10-jointed; elytra with punctate rows; proleg sexually monomorphic, protarsus not thickened in male. Oryctophileurus  and Amblyodus  are distinguished from the other mentioned genera by large horns, a distinctly developed and posteriorly carinate pronotal cavity and four external teeth on the protibia ( Endrödi 1985; Ide 1998). However, in Amblyodus  the two cephalic horns lack the lateral carina, the punctures in the pronotal cavity are predominately fused to short, transverse ridges, and the teeth close to the anterior pronotal margin are absent. Except for the horn number, these generic differences between Oryctophileurus  and Amblyodus  have not been recognized by previous workers.

Based on the combination of apomorphic characters of Oryctophileurus  that lacks in Amblyodus  we suggest retaining the genus Oryctophileurus  .

Key to species of Oryctophileurus