Ptomaphaginus scaphaner Szymczakowski, 1972

Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva, 2018, A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae), ZooKeys 777, pp. 57-108 : 75

Ptomaphaginus scaphaner Szymczakowski, 1972 Figures 5c, 7c, d, 9i, 10f

Ptomaphaginus scaphaner Szymczakowski, 1972: 279-300, figs 28-33); type from Cue phuong Ninh binh, Vietnam (in HNHM).

Material.

(in addition to that given in Schilthuizen and Perreau (2008)): Sabah. Kinabalu Park, Silau-Silau (low), 1530 m elev., human excrement traps, 7-11.xi.1987 (leg. Krikken & Rombaut, RMNH.INS.1086209-1086214), 5 males, 1 female; Kinabalu Park, Bukit Ular trail (low), 1800 m elev., human excrement traps, 7-11.xi.1987 (leg. Krikken & Rombaut, RMNH.INS.1086224-1086231), 3 males, 5 females; Kinabalu Park, Mempening trail (high), 1700 m elev., human excrement traps, 7-11.xi.1987 (leg. Krikken & Rombaut, RMNH.INS.1086233), 1 male; Crocker Range, Kota Kinabalu-Tambunan road (km 56), 1350 m elev., fish traps, 21-24.xi.1987 (leg. Krikken & Rombaut, RMNH.INS.1086215-1086223), 2 males, 7 females; Kinabalu Park, Headquarters Area, Liwagu Trail, 6°00'29.2"N 116°32'43.8"E, pitfall with chicken, 2.iv.2016 (leg. M. Schilthuizen, I. Njunjić & F. Feijen) 2 males (BORN), 1 male and 1 female (RMNH.INS.1086232). Sarawak. Mulu National Park, Limestone 6.4, Tp 113, 400 m elev., 6.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 male; Mulu National Park, Limestone 6.4, Tp 110, 650 m elev., 6.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 male, 2 females; Mulu National Park, mixed dipterocarp forest litter, TPS 5-6, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 4 males, 1 female; Mulu National Park, mixed dipterocarp forest litter, Tp 7, 21.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 male; Mulu National Park, slope, baited traps, 7.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 3 females; Mulu National Park, slope, TPS 4-6, 29.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 female; Mulu National Park, TPS 7-13, 4.v.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 female.

Description.

Length 1.8-2.5 mm. Habitus broad and short, colouration light reddish-brown. Pronotum 1.75 times as wide as long, slightly narrower than the elytra. Elytra 1.2 times as long as jointly wide (length measured from the caudal tip of the scutellum to the caudal tip of the elytra). Wings present. Antennae very broad and short, with antennomeres 9 and 10 twice as broad as long. Aedeagus strongly curved, extremely convex and swollen, ending in a flattened “beak”; stylet long and thin, hair-like, running along the inside of the roof of the convex part of the aedeagus. Spiculum gastrale short, triangular, as long as wide. Spermatheca U-shaped, with 6-7 broad rings along the proximal part, and ca. 5 much narrower rings in the terminal one-quarter. Spermiduct very long and very thin.

Differential diagnosis.

Unique among the Bornean Ptomaphaginus by the condensed antennae and the inflated aedeagus with flattened “beak” and long stylet.

Habitat and distribution.

This appears to be a very widespread species, now known from Vietnam, Peninsular Malaysia, Borneo, and Java. It is possible, however, that the species consists of a complex of closely related species, given the geographic variation in secondary sexual characters ( Schilthuizen and Perreau 2008).