Aposphragisma kolleri Thoma, 2014

THOMA, MARCO, KRANZ-BALTENSPERGER, YVONNE, KROPF, CHRISTIAN, GRABER, WERNER, NENTWIG, WOLFGANG & FRICK, HOLGER, 2014, <strong> The new Southeast Asian goblin spider genus <em> Aposphragisma </ em> (Araneae, Oonopidae): diversity and phylogeny </ strong>, Zootaxa 3798 (1) : 44-46

Aposphragisma kolleri Thoma , sp. nov.

( Figs. 26–27 )

Type material. HOLOTYPE: male (PBI_OON 00031373), Sarawak ( MALAYSIA), Bako National Park [01°42′N 110°27′E; inf.], 30.III.1985, lowland forest, litter, leg. C. Deeleman, deposited at RMNH . PARATYPE: 1 male (PBI_OON 00031386), collected together with holotype, deposited at RMNH .

Etymology. The species epithet is dedicated to Markus Koller-Furrer (*1958), a Swiss teacher, naturalist and conservationist who promoted the first author’s ornithological and natural history skills from an early age.

Diagnosis. A. kolleri sp. nov. belongs to the stripe-clade and can be distinguished from other species of this clade by the combination of the following traits: carapace with one pair of spikes (csp) ( Figs. 26 C–D); carapace margin with blunt denticles, sluice restricted to posterior margin ( Fig. 26D ). The new species resembles A. brunomanseri sp. nov. and A. dentatum sp. nov. but differs from A. brunomanseri sp. nov. among others by the lack of a second pair of spikes on the carapace, by the sternum shape and the pointed embolic spine (esp) ( Fig. 27C ), from A. dentatum sp. nov. among others by the shape of the carapace margin denticles, shorter claws (almost as long as tarsi in A. dentatum sp. nov.) and greater body length.

Description. Description based on 2 males.

MALE. With the characters of the genus except as noted. Body length 2.08–2.16 mm, carapace length 0.96 mm. Sclerotized parts uniformly coloured red-brown, approaching colour 32 ‘chestnut’; legs paler, orangish. Habitus: Figs. 26 A–C.

CEPHALOTHORAX. Carapace: surface of elevated portion of pars cephalica smooth, sides strongly reticulate ( Fig. 26D ); posterior edge of pars cephalica with one pair of spikes (csp) ( Figs. 26 C–D); carapace margin with sluice restricted to posterior margin, with blunt denticles ( Figs. 26 D–E). Eyes: posterior eye-row straight from above, procurved from front ( Figs. 26D, F ); ALE largest, separated by their radius to diameter; ALE-PLE separated by less than ALE radius; PME touching throughout most of their length; PLE-PME separated by less than PME radius. Sternum finely ornamented except smooth median stripe ( Fig. 26E ); posterior margin with narrow single extension, covered with blunt denticles ( Fig. 26E ; cf A. brunomanseri sp. nov.). Pleura: surface smooth with pairs of large pits dorsally of each coxa I, II and III. Mouthparts: chelicerae posterior margin of inner surface proximally modified to a ridge with median slit, covering about 2/3 of margin length, distally ending in a rounded bulge; promargin with row of flattened setae, distally extending into a short inwards pointing tooth-like projection (tlp) ( Fig. 26F ); posterior face with a pair of long, spatulate setae.

ABDOMEN. Book lung covers (bc) large, elongated, elliptical to rather dumbbell shaped (sides concave), about 4–5 times longer than wide ( Fig. 26G ). Epigastric scutum dorsally with widely oval subterminal ridge (sr) ( Fig. 26G ); scuto-pedicel region with slightly swollen hair bases. Postepigastric scutum long, semicircular, with long posteriorly directed lateral apodemes ( Fig. 26H ). Dorsal abdominal setae shorter than in A. brunomanseri sp. nov. ( Fig. 26G , cf fig. 9C).

GENITALIA. Epigastric region: sperm pore situated between anterior and posterior spiracles ( Fig. 26H ). Palp: basal segments as well as bulb and cymbium lighter in colour than rest of body; bulb stout ( Figs. 27 A–B). Embolus (em) medially with longitudinal ridges on prolateral surface, tip spatulate ( Fig. 27C ); embolic spine (esp) with pointed tip ( Fig. 27C ). Conductor (con) with broad tip (appearing ‘snout’-like in lateral view; Fig. 27C ). Embolusconductor-complex as long as bulb.

Intraspecific variation. Body size variable (see above); male paratype slightly paler and with slightly longer spikes on carapace.

Distribution. W-Sarawak, MALAYSIA ( Fig. 49 ).

Figures

FIGURE 26. Aposphragisma kolleri sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace front view (left fang artificially displaced); G. abdomen anterolateral view; H: abdomen ventral view. Bc, book lung cover; csp, posterior cephalic spikes (modified hair bases); ctg, continuous transverse groove; lap, postepigastric lateral apodemes; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–F, H 500 µm; G 200 µm.

FIGURE 27. Aposphragisma kolleri sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view. Cb, conical bulb; con, conductor; em, embolus; esp, embolic spine; spp, spermophor. Scale bars: A–B 100 µm; C 50 µm.

FIGURE 49. Distribution of Aposphragisma gen. nov. in Southeast Asia. Stripe clade: 1) A. baltenspergerae sp. nov., 2) A. borgulai sp. nov., 3) A. brunomanseri sp. nov., 4) A. dayak sp. nov., 5) A. dentatum sp. nov., 6) A. hausammannae sp. nov., 7) A. helvetiorum sp. nov., 8) A. kolleri sp. nov., 9) A. menzi sp. nov., 10) A. monoceros sp. nov., 11) A. retifer sp. nov., 12) A. salewskii sp. nov., 13) A. sepilok sp. nov. Non-stripe clade species: 14) A. confluens sp. nov., 15) A. draconigenum sp. nov., 16) A. nocturnum sp. nov., 17) A. rimba sp. nov., 18) A. scimitar sp. nov., 19) A. stannum sp. nov. Scale bars: 500 km.