Cloeodes magnus Salles, Massariol & Angeli
Salles, F. F., Massariol, F. C., Angeli, K. B., Lima, M. M., Gattolliat, L. & Sartori, M., 2015, Revealing the diversity of Cloeodes Traver, 1938 (Ephemeroptera: Baetidae) in the Neotropics: description of eleven new species from Brazilian mountain ranges, Zootaxa 4020 (1), pp. 1-50 : 31
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|Cloeodes magnus Salles, Massariol & Angeli|
Cloeodes magnus Salles, Massariol & Angeli , sp. nov.
Figures 2 View FIGURE 2 d, 21, 22, 30.
Diagnoses. NYMPH. A) Large body size (> 7.0 mm); B) Body color pattern with distinct darker marks on abdominal terga ( Fig. 2 View FIGURE 2 d); C) Labrum, medially, with two long setae ( Fig. 21 View FIGURE 21 a); D) Maxillary palp short (¾ length of galea-lacinia, Fig. 21 View FIGURE 21 d); E) Inner margin of labial palp segment III subequal to margin ( Fig. 21 View FIGURE 21 e); F) Distal projection of fore femur short (not exceeding apex of femur, Figs. 22 View FIGURE 22 a, 22 b); G) Gills without medial rib ( Fig. 22 View FIGURE 22 e).
Nymph (nearly mature). Lengths. Body: 10.4 –12.0 mm; cerci: 3.9 mm; terminal filament: 3.9 mm; antenna: 3.0.
Coloration ( Fig. 2 View FIGURE 2 d). General coloration yellowish-white profusely washed with dark brown. Head yellowish washed with brown, except around lateral ocellus and below median ocellus. Legs yellowish washed with brown, except for two unpigmented stripes close to inner margin of femur; tarsus darker. Terga. Segments IV and VIII less pigmented; segment IX with a medial less pigmented area; segments II, III and VII sometimes less pigmented, but never as in segments IV and VIII; segments I–IX medially with a longitudinal unpigmented stripe. Sterna yellowish washed with brown, darker on segments VII–IX; segments III–IX with a sublateral brown stripe. Filaments yellowish-white washed with brown on basal half and washed with dark brown on apical ¼.
Head. Labrum ( Fig. 21 View FIGURE 21 a). Length about 0.6 × maximum width; dorsal arc with 2 medial setae; ventrally with 5–6 spine-like setae near lateral margin.
Mandibles ( Figs. 21 View FIGURE 21 b, 21 c). Denticles of mola constricted at base (as in fig. 6 f).
Maxilla ( Fig. 21 View FIGURE 21 d). Medially with 1 spine-like seta + 9 apically pointed setae. Maxillary palp short (¾ length of galea-lacinia, Fig. 21 View FIGURE 21 d); palp segment II 2.0 × length of segment I.
Labium ( Fig. 21 View FIGURE 21 e). Base of glossa expanded (reaching at least half of paraglossa); inner and outer margin respectively with around 20 and 15 spine-like setae increasing in length apically; ventral surface scattered with few short setae at base. Paraglossa dorsally with arc of 3 setae close to inner margin; ventrally with arc of 8–10 setae close to inner margin. Labial palp with segment I 0.9 × length of segments II and III combined; segment II dorsally with oblique row of 5 setae; segment III with distal margin slightly truncate; inner margin subequal to outer margin; ventrally covered with around 30 long setae ( Fig. 21 View FIGURE 21 e); setae on inner margin serrate.
Thorax. Hind wing pad. Present.
Fore leg ( Fig. 22 View FIGURE 22 a). Ratio 1.8: 1 (0.8mm): 1: 0.2. Femur. Length about 4.5 × maximum width; outer margin with row of 11–12 blunt, clavate setae (length of setae about 0.1 × maximum width of femur); submarginal setae absent; anterior surface and inner margin with around 40 spine-like setae; posterior surface bare; distal projection distinct, but not exceeding apex of femur ( Fig. 22 View FIGURE 22 b), 0.1 × width of femur, apically with 3 clavate setae. Tibia. Outer margin with many simple setae; inner margin with one simple seta on apex; subtending bristle blunt, clavate, extending apex of tibia ( Fig. 22 View FIGURE 22 c). Tarsus. Inner margin with row of 14 + 1 spine-like setae. Tarsal claw. 0.3 × length of tarsus.
Mid and hind legs. Similar to fore leg, except for the lower number of setae on anterior surface of femur, subapical projections of femur less developed and type of setae on inner apex of tibia (simple instead of serrate).
Abdomen. Terga ( Fig. 22 View FIGURE 22 d). With spines present on posterior margin of segments I–X; large spines intercalated by short spines, larger spines 4.9 × longer than wide.
Sterna . With spines present on posterior margin of segments IV–IX.
Gills ( Fig. 22 View FIGURE 22 e). Outer margin serrated on distal 1 / 4; inner margin slightly serrated; medial rib absent; tracheae extending from main trunk to inner and outer margins. Gill I subequal in length to segment II, oval. Gill IV reaching 1 / 2 of segment VI, asymmetric, broad at middle. Gill VII reaching 1 / 2 of segment IX.
Paraproct ( Fig. 22 View FIGURE 22 f). With 12–14 spines.
Cercus . With large spines on every four segments at base and on every two segments toward apex.
Terminal filament. With spines on every four segments.
Male and female imago. Unknown.
Distribution ( Fig. 30 View FIGURE 30 ). Southern Brazil, Santa Catarina State, Serra Geral.
Type-material. HOLOTYPE. Nymph (on two slides, one with mouth parts, legs and gills, one with abdomen), BRAZIL, Santa Catarina, Urubici, Cachoeira do Avencal, S 28 °02’ 55.1 ” W 49 ° 37 ’00.0”, 1220 m.a.s.l., 02/x/ 2011, Boldrini R, col. ( CZNC). PARATYPES. 5 nymphs, same data as holotype (3 CZNC, 2 INPA).
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