Anomalophylla subcarinata, Ahrens, 2005

Ahrens, Dirk, 2005, Taxonomic revision of the genus Anomalophylla Reitter, 1887 (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), Zootaxa 1076 (1), pp. 1-62 : 18-20

Anomalophylla subcarinata sp. n.

( Fig. 1N–P , 6 )

Type material. Holotype: ” China Cent. Sichuan Luding Co. Moxi env. 22.5.–10.6.93

Benes” ( ZSM). Paratypes : 2 ♂♂, 7 ♀♀ same data as holotype ( CA, TICB), 1 ♂, 1 ♀

“ CHINA, SW Sichuan Moximian, 1300 m Krajcik M. lgt. 1.6.97” ( CA), 2 ♂♂, 2 ♀♀ “Ta­

tsien­Lou Chasseurs Thibétains 1896 / Muséum Paris 1952 Coll. R. Oberthür” ( MNHN), 2

♂♂ “Thibet Chasseurs de Ta­tsien­Lou 1895 / Muséum Paris 1952 Coll. R. Oberthźr

( MNHN), 1 ♂ “N. YUNNAN 19–25.VI.1994 27°49’N 99°43’E cca 3600 m ZHONGDIAN lgt. D. Kr á l & J. Farka č ( NHMB), 7 ♂♂, 1 ♀ “W SICHUAN 1–7.VII.1994 29°37’N 102°07’ 1200–1900 m MOXI­HAILOUGOU lgt. J. Farka č & D. Kr á l ( NHMB,

CA, CK).

Holotype description. Length: 5.7 mm, length of elytra: 3.6 mm, width: 3.1 mm. Body oblong, black, dorsal surface dull and with long, dense, erect setae; pilosity black, setae on elytra and sometimes those on pronotum posteriorly white. Head: Labroclypeus transverse, widest medially; lateral margins strongly convex and convergent anteriorly; anterior angles strongly rounded; lateral border and ocular canthus producing a distinct blunt angle, anterior and lateral margin strongly reflexed, anteriorly moderately sinuate medially; surface almost flat and moderately shiny, coarsely and densely punctate with numerous long, erect setae. Frontoclypeal suture distinct, weakly curved and slightly elevated; smooth area anterior to eye as wide as long. Ocular canthus moderately long and slender, finely and densely punctate, densely setose. Frons with coarse, dense punctures; punctures bearing a long, erect seta; basal punctation less dense, more irregular. Eyes small, ratio of diameter / interocular width: 0.48. Antenna dark yellow, with ten antennomeres; club black with five equal in length antennomeres; club 2.5 times as long as the remaining antennomeres combined, reflexed. Prementum almost flat. Pronotum: widest at base; lateral margins in posterior half almost straight and subparallel, in anterior half strongly convex and convergent; anterior angles weakly produced and moderately rounded, posterior angles blunt and moderately rounded; anterior margin medially weakly convex with distinct, fine marginal line; basal margin with a fine marginal line. Pronotal surface with dense, double punctation; fine glabrous punctures mixed with large punctures bearing a long erect seta which is slightly directed anteriorly and weakly curved posteriorly; anterior and lateral borders setose; basal margin of hypomeron not produced ventrally, not transversely sulcate anterior to base. Scutellum : moderately long, apex weakly rounded with fine, dense punctures; posteriorly partially smooth; minute setae present in the punctures. Elytra: oblong, widest medially, striae indistinctly impressed and finely densely punctate. Elytral intervals flat with fine, moderately dense punctures; with long, erect setae in the punctures on all intervals; even intervals anteriorly setose only; sutural interval with robust, long single seta. Epipleural edge fine, ending at the strongly convex external apical angle of elytra, epipleura densely setose, strongly curved in anterior third of elytra; apical border chitinous, without short microtrichomes. Venter: Ventral surface dull with fine, dense punctures; metasternum with dense, long setae; setae partially appressed, partially erect. Metacoxa with fine, long, appressed setae; glabrous medially. Abdominal sternites with indistinct transverse row of coarse punctures bearing thick setae between fine, dense punctation; all sternites with fine, long setae; tegument of abdominal sternites (60x magnification) with fine polygonal mesh pattern formed by microtrichomes; ultimate three abdominal sternites at midline with longitudinally impressed line. Mesosternum between mesocoxae narrow, narrower than mesofemur; with irregularly scattered, strong setae. Ratio of length of metepisternum / metacoxa: 1 / 1.43. Pygidium moderately convex, completely shiny; with fine, dense punctures bearing fine, long setae; without smooth midline. Legs: slender with shiny surface. Femora with two longitudinal rows of setae, finely densely punctate and setose; anterior edge of metafemur acute, lacking adjacent serrated line; posterior margin weakly convex with a few fine setae medially, ventral posterior margin weakly widened in apical half; posterior margin smooth ventrally and dorsally. Metatibia moderately slender, long, widest at apex; ratio width / length: 1 / 3.5; dorsal margin sharply carinate and smooth with two groups of spines, the basal group at one third, apical one at two thirds of metatibial length. Metatibia basally with a few single spines in punctures; lateral face longitudinally convex with moderately dense, fine punctures; ventral edge serrated, with four strong, long, equally spaced spines; medial face finely punctate, apex sharply truncate interiorly near tarsal articulation. Tarsomeres dorsally glabrous, coarsely punctate; ventrally with sparse, short setae; metatarsomeres dorsally with longitudinally impressed punctures, ventrally with strongly serrated ridge, laterally without strong longitudinal carina; first metatarsomere slightly shorter than the following two tarsomeres combined and one third of its length longer than the upper tibial spur. Protibia short, bidentate. Protarsal claws symmetrical, basal tooth of inner protarsal claw evenly pointed. Aedeagus: Fig. 1N–P .

Intraspecific variation. Length: 5.7–6.7 mm, length of elytra: 3.4–4.1, width: 3.0– 3.4 mm. Some specimens have the elytra uniformly reddish brown or dark brown. The density of punctation on dorsal surface as well as shape of lateral border of pronotum are variable. ♀: Antennal club with three antennomeres and distinctly shorter than remaining antennomeres combined; eyes equal in size to that of male.

Diagnosis. The new species is similar in external habitus as well as in genital morphology to A. tristicula . It differs by the sharply pointed right paramere and its shorter and sharply pointed basal lobe. The lateral phallobasal apophysis of A. subcarinata is narrower and more strongly pointed.

Etymology. Combination of the Latin prefix sub ­ (fairly) and adjective carinatus (carinate), intending weakly carinate.

Figures

FIGURE 1. (A–C) Anomalophylla tristicula Reitter (Tibet: Kuku­Nor); (D–F) A. tristicula Reitter (China, Yunnan, Dequen env., “population B”); (G–J) A. tsangpoana sp. n. (holotype); (K–M) A. moupinea (Fairmaire) (lectotype “Chine: Moupin”); (N–P) A. subcarinata sp. n. (holotype); (Q–S) A. obscuripennis sp. n. (holotype). (A, D, G, K, N, Q) aedeagus, left side lateral view; (B, E, H, L, O, R) parameres, dorsal view; (C, F, J, M, P, S) aedeagus, right side lateral view. Scale bar = 0.5 mm.

FIGURE 6. Present known distribution of Anomalophylla liciata, A. moupinea, A. subcarinata, and A. subfastuosa.