Bembidion (Nothonepha), Jeannel, 1962

Maddison, David R., 2014, An unexpected clade of South American ground beetles (Coleoptera, Carabidae, Bembidion), ZooKeys 416, pp. 113-155 : 132

Taxon classification Animalia Coleoptera Carabidae

Subgenus Nothonepha Jeannel, 1962

Nothonepha Jeannel, 1962; type species Bembidium baptisatum Csiki (= Bembidion lonae Jensen-Haarup), by original designation.

Remarks.

As here defined, the subgenus Nothonepha includes all species of the Antiperyphanes Complex possessing mesepisternal pits. Seven described species belong to Nothonepha :

Bembidion lonae Jensen-Haarup, 1910 (Fig. 5D)

Bembidion pallideguttula Jensen-Haarup, 1910

Bembidion eburneonigrum Germain, 1906 (Fig. 5E)

Bembidion engelhardti Jensen-Haarup, 1910

Bembidion engelhardti engelhardti Jensen-Haarup, 1910 (Fig. 5G)

Bembidion engelhardti rayoda Toledano, 2008

Bembidion tucumanum (Jeannel, 1962) (Fig. 5F)

Bembidion germainianum Toledano, 2002 (Fig. 5C)

Bembidion tetrapholeon Maddison, sp. n. (Figs 5A, B)

Four of these species ( Bembidion germainianum , Bembidion tucumanum , Bembidion engelhardti , and Bembidion eburneonigrum ) were formerly placed in subgenus Antiperyphus . In addition, there are at least three undescribed species (including Bembidion sp. nr. lonae, sequenced here). The species figured by Toledano (2008) as Bembidion germainianum is an undescribed species related to Bembidion germainianum . A revision of the subgenus is in preparation ( Roig-Juñent and Maddison).