Ummidia huascazaloya, Godwin & Bond, 2021

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia huascazaloya sp. nov. Figs 40 , 41 , Map 5

Type material.

HOLOTYPE ♂ (CNAN-T01384) from just outside Peña del Aire Park, outside Huasca de Ocampo, Hidalgo, Mexico, 20.2762 -98.5169 1, 2046 m a.s.l., 6-7.vi.2019, coll. PE Cushing, H Carmona, DL Batista Perales, J Rojas Castillo, CNAN. PARATYPE: 1 ♀ (UMM171) from Chapulhuacán, Hidalgo, Mexico, 21.1569 -98.9026 6, 967 m a.s.l.. coll. F Bonet, AMNH.

Etymology.

The specific epithet is a noun taken in apposition and is in reference to the original Nahuatl name for Huasca de Ocampo.

Diagnosis.

Ummidia huascazaloya can be differentiated from U. anaya by the presence of a defined comb on the retrolateral face of tarsus IV in males and females. U. huascazaloya males can be further differentiated from U. anaya by having a palpal tibia (3.25 × long as wide vs 1.85 ×) and tarsus IV (3.6 × long as wide vs 2.9 ×) which are more elongate as well as having fewer, larger endite cuspules (10 vs 22). Males can be differentiated from U. cuicatec and U. zilchi by having a sinuous embolus without a strong bend and from U. cuicatec , U. zebrina , U. brandicarlileae , and U. yojoa by having more spines on the prolateral face of tibia I (11 vs 0-2). Males can be differentiated from U. riverai by lacking a pale dorsal heart patch on the abdomen. Females can be differentiated from U. gabrieli , U. pesiou , U. rodeo , U. pustulosa , and U. frankellerae by having spermathecae that bend medially then anteriorly. Males disperse in June.

Description of male holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Right and left leg III missing. Cephalothorax. Carapace 7.36 long, 7.12 wide. Pars cephalica 5.25 long. Foveal groove procurved, 0.48 long, 1.76 wide. Tubercle moderate, mostly restricted to ME. AER procurved. PER recurved. Eye group 0.88 long, 1.62 wide, AME 0.34, PME 0.24, ALE 0.51, PLE 0.3. Sternum moderately setose, STRl 4.31, STRw 3.88. Chelicerae with anterior tooth row comprising six teeth, posterior margin with six teeth. Palpal endites with ten relatively long non-haste spinules over proximal half of endite face, lacking distal endite cuspules, ENDw 1.59, ENDl 2.72. Labium with three relatively long, non-hastate cuspules, LBw 1.44, LBl 1.12. Rastellum with many spines on process. Abdomen setose with pale speckles concentrated at apodemes. Legs. F1 6.56; F1w 1.68; P1 3.12; Ti1 4.4; Mt1 3.2; Tr1 1.52; F4 6.48; F4w 1.96; P4 2.88; Ti4 3.92; Mt4 4.4; Tr4 2.24. Retrolateral face of tarsus IV with defined comb with heavier spinules over distal half. Leg I spination pattern: TSp 6, TSpv 5, TSrd 0, TSr 1, TSrv 17, MtSp 3, MtSr 12, TrSp 2, TrSr 8. Pedipalps. PTl 3.9, PTw 1.2, Bl 3.7. Embolus long; sinuous, bulb relatively small.

Variation, males.

Known only from male holotype specimen.

Description of female paratype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. General coloration. Carapace, chelicerae, and legs dark reddish brown 2.5YR 2.5/4, Tr IV strong brown 7.5YR 5/8. Abdomen black 7.5YR 2.5/1, spinnerets yellowish brown 10YR 5/6. Cephalothorax. Carapace 4.99 long, 4.71 wide. Pars cephalica 3.69 long. Foveal groove procurved, 1.09 long, 0.47 wide. All eyes on moderate tubercle. AER procurved. PER straight. Eye group 0.78 long, 1.22 wide, AME 0.3, PME 0.3, ALE 0.41, PLE 0.34. Sternum very sparsely setose around edges, thicker anteriorly STRl 3.03, STRw 2.97. Chelicerae with anterior row comprising five teeth, posterior margin with six teeth. Palpal endites with 30 very large cuspules spread across proximal half of endite face and 15 smaller cuspules distally, ENDw 1.23, ENDl 2.09. Labium with 13 very large cuspules, LBw 1.19, LBl 0.91. Rastellum with large blunt spines along rastellar margin and several smaller blunt spines on rastellar process. Abdomen. Evenly setose with pale speckles. Legs. F1 3.36; F1w 1.27; P1 2.09; Ti1 1.98; Mt1 1.33; Tr1 1.72; F3 2.86; F3w 1.57; P3 1.67; Ti3 1.61, Sd3 1.09; Mt3 1.24; Tr3 1.12; F4 3.59; F4w 1.5; P4 2.04; Ti4 1.92; Mt4 1.9; Tr4 1.02. Retrolateral face tarsus IV with comb of alternating long and short spinules. Pedipalps. PF 3.25, PP 1.72, PTi 1.81, PTr 1.67. Spermathecae bent medially then folded anteriorly, bulbs facing anteriorly.

Variation, females.

Known only from female paratype specimen.

Known only from type material.

Figures

Map 5. Locality records for southern Mexico and Central American species.

Figure 40. Ummidia huascazaloya sp. nov. from Hidalgo, Mexico A male habitus illustration CNAN-T 01384 B female habitus illustration UMM 171. Scale bars: 4.0 mm.

Figure 41. Ummidia huascazaloya sp. nov. from Hidalgo, Mexico A-D male holotype (CNAN-T 01384) A retrolateral aspect, leg I B prolateral aspect, leg I C line drawings, leg I retrolateral and prolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female paratype (UMM 171). Scale bars: 1.0 mm.