Exocelina hintelmannae (Shaverdo, Sagata & Balke, 2005)

Shaverdo, Helena, Sagata, Katayo & Balke, Michael, 2016, Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 577, pp. 125-148 : 128-130

Taxon classification Animalia Coleoptera Dytiscidae

2. Exocelina hintelmannae (Shaverdo, Sagata & Balke, 2005) Figs 4, 9F, G

Papuadytes hintelmannae Shaverdo, Sagata & Balke, 2005: 272.

Exocelina hintelmannae (Shaverdo, Sagata & Balke, 2005): Nilsson 2007: 33 (comb. n.); Toussaint et al. 2014: Supplementary figs 1-4, tab. 2. (MB 1367).

Type locality.

Papua New Guinea: border Simbu and Eastern Highlands Provinces, Crater Mountain, between Wara Sera Station and Herowana Village, Hulene River, approximately 06°43.4'S; 145°05.6'E.

Type material studied.

Holotype: male "264 DNA M Balke" [green], "PNG Simbu / EHPr. Crater Mountain, Sera - Herowana, Wara Hulene, 1000 m, 16IX2002, Balke & Sagata (PNG 17)", "Holotypus Papuadytes hintelmannae sp.n. des. H. Shaverdo, K. Sagata & M. Balke, 2005" (BMNH). Paratypes: 1 male "256 DNA M Balke" [green], "Papua New Guinea Simbu / EHPr. Crater Mountain, Wara Sera Station, 800 m, 14IX2002, Balke & Sagata (PNG 10)", "Paratypus Papuadytes hintelmannae sp.n. des. H. Shaverdo, K. Sagata & M. Balke, 2005" (NHMW). 1 male "260 DNA M Balke" [green], "PNG Simbu / EHPr. Crater Mountain, Sera - Herowana, upper Oh River, 1200 m, 15IX2002, Balke & Sagata (PNG 12)", "Paratypus Papuadytes hintelmannae sp.n. des. H. Shaverdo, K. Sagata & M. Balke, 2005" (NHMW).

Additional material.

Simbu/EHL: 1 female "Papua New Guinea: Simbu / EHPr. Crater Mountain, Wara Sera Station, 820m, 14IX2002, Balke & Sagata, (PNG 8)" (ZSM). 2 males, 6 females "Papua New Guinea: Simbu / EHP. Crater Mountain, Wara Sera Station, 800m, 14IX2002, Balke & Sagata, (PNG 009)" (ZSM). 17 males, 15 females "Papua New Guinea: Crater Mountain, Wara Sera Station, 800m, 14IX2002, Balke & Sagata, (PNG 010)" (NARI, NHMW, ZSM). 8 males, 9 females "Papua New Guinea Simbu/EHP. Crater Mountain, Wara Sera Station, 800m, 14IX2002, Balke & Sagata, (PNG 10)" (NHMW, ZSM). 3 males, 4 females "Papua New Guinea: Simbu / EHP. Crater Mountain, Sera - Herowana, Wara Pima, 900m, 15IX2002, Balke & Sagata (PNG 011)" (ZSM). 7 males, 14 females "Papua New Guinea: Crater Mountain, Sera - Herowana, upper Oh river, 1200m, 15IX2002, Balke & Sagata (PNG 012)" (NHMW, ZSM). 1 male, 1 female "Papua New Guinea: Crater Mountain, Sera - Herowana, Jau river, 1100m, 15IX2002, Balke & Sagata (PNG 013)" (ZSM). 8 males, 7 females "Papua New Guinea: Crater Mountain, Sera - Herowana, Jau river, 1000m, 15IX2002, Balke & Sagata (PNG 015)" (NHMW, ZSM). 12 males, 10 females "Papua New Guinea: Simbu/EHP. Crater Mountain, Sera - Herowana, Hulene river, 1000m, 16IX2002, Balke & Sagata (PNG 017)" (NHMW, ZSM). 25 males, 33 females "Papua New Guinea: Simbu / EHP. Crater Mountain, Herowana, Yawasa River, 1200m, 17.IX.2003, Balke & Sagata (PNG 019)" (NARI, NHMW, ZSM). Simbu: 8 males, 13 females "Papua New Guinea: Supa Haia, 1032, 10.xi.2002, K. Sagata, (WB1)" (NHMW, ZSM). EHL: 1 male "Stn. No. 177a", "NEW GUINEA: E. Highland Dist., Wanatabe Valley. Nr. Okapa, c. 5000 ft. 5.ii.1965", "M.E. Bacchus. B.M. 195-120" (BMNH). Gulf: 30 males, 30 females "Papua New Guinea: Gulf, Marawaka, nr. Ande, 1000m, 10.xi.2006, 07.03.598S 145.4.375E, Balke & Kinibel (PNG 89)", one of them an additional green label "DNA M.Balke 1367" (NHMW, ZSM). 32 males, 35 females "Papua New Guinea: Gulf Province, Marawaka, Mala, 1400m, 11.xi.2006, 07.05.664S 145.44.467E, Balke & Kinibel, (PNG 90)" (NARI, NHMW, ZSM). 16 male, 12 females "Papua New Guinea: Gulf, Marawaka, Andakombe towards Morobe, 1100m, 12.xi.200, 07.09.766S 145.46.333E, Balke & Kinibel (PNG 92)" (NHMW, ZSM). 1 male, 3 females "Papua New Guinea: Gulf, Marawaka, Andakombe towards Morobe, 1500m, 12.xi.200, 07.10.413S 145.49.555E, Balke & Kinibel (PNG 93)", the male with an additional green label "DNA M.Balke 1362" (ZSM).

Additions to the description

(original description in Shaverdo et al. 2005, p. 272). Size and shape: Beetles medium-sized (TL-H 3.4-4.15 mm, TL 3.75-4.55 mm, MW 1.8-2.2 mm; holotype: TL-H 3.9 mm, TL 4.3 mm, MW 2.05 mm), see Fig. 4.

Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.

Variability: Beetles vary in shape of the median lobe and its ventral sclerite: mainly shape and length of its left lobe (Fig. 9F, G).

Distribution.

Papua New Guinea: Simbu, Eastern Highlands, and Gulf Provinces (Fig. 14).