Nebria (Sadonebria) quinquelobata, Sasakawa, Koji, 2016

Sasakawa, Koji, 2016, Two new species of the ground beetle subgenus Sadonebria Ledoux & Roux, 2005 (Coleoptera, Carabidae, Nebria) from Japan and first description of larvae of the subgenus, ZooKeys 578, pp. 97-113 : 100

Taxon classification Animalia Coleoptera Carabidae

Nebria (Sadonebria) quinquelobata sp. n. Figs 4, 8

Nebria saeviens (part): Uéno (1985): 56, fig. 11.

Nebria (Orientonebria) saeviens (part): Farkač and Janata (2003): 94.

Nebria (Sadonebria) saeviens (part): Ledoux and Roux (2005): 824, fig. 621; Yoshitake et al. (2011): 38.

Type materials.

Holotype: ♂ (NIAES), " X. 13, 1965 / Mt. Myoko / Niigata P. / K. BABA" [type locality: Mt. Myôkô, Myôkô-shi, Niigata Prefecture, Japan]. Paratypes: 2♂ 1♀ (NIAES), "Sasagamine [in Japanese] / S-Echigo / 28.VIII, 1966 / Col. K. Baba".

Etymology.

The specific name derives from the Latin adjectives quinque- (five of) and lobatus, -a, -um (with lobes) and refers to the ventral view of the male endophallus (Fig. 8c).

Diagnosis.

Similar to Nebria saeviens (Fig. 5) in having yellowish brown legs but distinguished by the shape of the endophallus (Figs 8, 9).

Description.

External structures: Body length (including mandibles): ♂, 11.7-12.3 mm (mean ± SD: 12.0 ± 0.30 mm, n = 3); ♀, 13.1 mm (n = 1). Head with a reddish-brown patch between the eyes; pronotum reddish brown on the central part and dark brown at the lateral margins; head and other pronotal parts and elytra black; legs yellowish to light brown.

Male genitalia: Laterobasal lobes and lateroapical lobes both largely swollen and widely rounded, with the former slightly smaller than the latter; the dorsoapical lobe similar in size to the laterobasal lobe and bifurcated in a T shape at the apex; dorsobasal lobe distinct but smaller than the other lobes.